.......................... ooooooooooooooooooooooooooooo CURRENT SETTINGS ooooooooooooooooooooooooooooooooooo .......................... . Sequence filename : nucleic_M2564_142x1130_2006.phy . Data type : dna . Sequence format : interleaved . Number of data sets : 1 . Nb of bootstrapped data sets : 0 . Model name : GTR . Proportion of invariable sites : 0.000000 . Number of subst. rate categs : 4 . Gamma distribution parameter : estimated . 'Middle' of each rate class : mean . Nucleotide equilibrium frequencies : empirical . Optimise tree topology : yes . Tree topology search : Best of NNIs and SPRs . Starting tree : BioNJ . Add random input tree : yes . Number of random starting trees : 5 . Optimise branch lengths : yes . Optimise substitution model parameters : yes . Run ID : none oooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooo . Compressing sequences... . 780 patterns found. (out of a total of 1130 sites) . 486 sites without polymorphism (43.01%). . [Random start 1/ 5] . Computing pairwise distances... . Building BioNJ tree... . Randomising the tree... . This analysis requires at least 59Mo of memory space. . Maximizing likelihood (using SPR moves)... . ( 6 sec) [ -43993.2965] [GTR parameters ] . ( 7 sec) [ -42984.2445] [Alpha ][ 0.500960] . ( 7 sec) [ -42872.5170] [ 7429] [depth= 7] . ( 8 sec) [ -42788.7616] [ 7413] [depth= 9] . ( 8 sec) [ -42709.4102] [ 7404] [depth= 1] . ( 9 sec) [ -42664.9899] [ 7393] [depth= 9] . ( 9 sec) [ -42612.9902] [ 7386] [depth= 1] . ( 9 sec) [ -42545.9623] [ 7378] [depth= 15] . ( 9 sec) [ -42483.7551] [ 7378] [depth= 7] . ( 9 sec) [ -42477.1417] [ 7377] [depth= 5] . ( 10 sec) [ -42458.9084] [ 7374] [depth= 3] . ( 10 sec) [ -42412.2089] [ 7373] [depth= 7] . ( 10 sec) [ -42380.6113] [ 7368] [depth= 5] . ( 10 sec) [ -42297.0479] [ 7367] [depth= 12] . ( 11 sec) [ -42286.4478] [ 7360] [depth= 5] . ( 11 sec) [ -42267.2539] [ 7358] [depth= 13] . ( 11 sec) [ -42260.2079] [ 7358] [depth= 2] . ( 11 sec) [ -42259.6913] [ 7357] [depth= 1] . ( 12 sec) [ -42210.9184] [ 7354] [depth= 5] . ( 12 sec) [ -41966.0815] [ 7353] [depth= 2] . ( 12 sec) [ -41931.5818] [ 7351] [depth= 4] . ( 12 sec) [ -41880.5270] [ 7351] [depth= 1] . ( 13 sec) [ -41877.9082] [ 7351] [depth= 1] . ( 13 sec) [ -41872.0411] [ 7353] [depth= 1] . ( 13 sec) [ -41853.8554] [ 7349] [depth= 2] . ( 13 sec) [ -41710.4511] [ 7334] [depth= 14] . ( 14 sec) [ -41639.2786] [ 7319] [depth= 12] . ( 14 sec) [ -41634.5293] [ 7318] [depth= 13] . ( 14 sec) [ -41596.2267] [ 7316] [depth= 18] . ( 15 sec) [ -41565.8560] [ 7309] [depth= 2] . ( 15 sec) [ -41530.1184] [ 7307] [depth= 7] . ( 15 sec) [ -41275.4007] [ 7293] [depth= 26] . ( 15 sec) [ -41036.4229] [ 7291] [depth= 1] . ( 16 sec) [ -40840.0152] [ 7225] [depth= 19] . ( 16 sec) [ -40533.6452] [ 7183] [depth= 17] . ( 16 sec) [ -40400.7016] [ 7142] [depth= 20] . ( 17 sec) [ -40210.1153] [ 7105] [depth= 17] . ( 17 sec) [ -40172.3098] [ 7100] [depth= 11] . ( 17 sec) [ -40141.7422] [ 7099] [depth= 1] . ( 17 sec) [ -39980.2610] [ 7062] [depth= 17] . ( 17 sec) [ -39555.3524] [ 7012] [depth= 15] . ( 18 sec) [ -39424.8664] [ 7000] [depth= 11] . ( 18 sec) [ -39369.9211] [ 6984] [depth= 11] . ( 18 sec) [ -39299.1110] [ 6981] [depth= 3] . ( 19 sec) [ -39224.7087] [ 6972] [depth= 6] . ( 19 sec) [ -39206.5022] [ 6971] [depth= 2] . ( 19 sec) [ -39184.3326] [ 6971] [depth= 1] . ( 19 sec) [ -39170.4368] [ 6967] [depth= 1] . ( 20 sec) [ -39001.8255] [ 6951] [depth= 30] . ( 20 sec) [ -38925.8261] [ 6886] [depth= 30] . ( 20 sec) [ -38912.6032] [ 6881] [depth= 1] . ( 20 sec) [ -38866.1737] [ 6874] [depth= 9] . ( 21 sec) [ -38769.5321] [ 6845] [depth= 12] . ( 21 sec) [ -38754.2335] [ 6846] [depth= 1] . ( 22 sec) [ -38702.2206] [ 6839] [depth= 26] . ( 22 sec) [ -38430.2276] [ 6827] [depth= 2] . ( 22 sec) [ -38056.0723] [ 6823] [depth= 25] . ( 22 sec) [ -37516.7661] [ 6728] [depth= 26] . ( 22 sec) [ -37329.8219] [ 6711] [depth= 2] . ( 23 sec) [ -37237.7753] [ 6711] [depth= 4] . ( 23 sec) [ -37042.2416] [ 6641] [depth= 11] . ( 24 sec) [ -36946.4433] [ 6630] [depth= 5] . ( 24 sec) [ -36901.4919] [ 6627] [depth= 15] . ( 24 sec) [ -36833.4091] [ 6611] [depth= 6] . ( 24 sec) [ -36746.1815] [ 6606] [depth= 1] . ( 25 sec) [ -36538.0680] [ 6559] [depth= 24] . ( 25 sec) [ -36525.0835] [ 6558] [depth= 2] . ( 25 sec) [ -36500.2039] [ 6553] [depth= 15] . ( 26 sec) [ -36449.3759] [ 6553] [depth= 2] . ( 26 sec) [ -36422.5280] [ 6553] [depth= 1] . ( 26 sec) [ -36406.2966] [ 6549] [depth= 3] . ( 26 sec) [ -36406.0089] [ 6548] [depth= 15] . ( 26 sec) [ -36304.9524] [ 6515] [depth= 17] . ( 27 sec) [ -36279.5309] [ 6506] [depth= 16] . ( 27 sec) [ -36212.4953] [ 6497] [depth= 2] . ( 27 sec) [ -36199.2524] [ 6491] [depth= 12] . ( 28 sec) [ -36138.8875] [ 6485] [depth= 12] . ( 28 sec) [ -35718.4956] [ 6454] [depth= 14] . ( 28 sec) [ -35717.2964] [ 6453] [depth= 1] . ( 28 sec) [ -35708.6251] [ 6450] [depth= 11] . ( 29 sec) [ -35336.5890] [ 6446] [depth= 13] . ( 29 sec) [ -35321.6291] [ 6443] [depth= 17] . ( 29 sec) [ -35246.8468] [ 6427] [depth= 18] . ( 29 sec) [ -35189.4230] [ 6419] [depth= 10] . ( 30 sec) [ -35185.2264] [ 6417] [depth= 1] . ( 30 sec) [ -35150.1155] [ 6416] [depth= 10] . ( 30 sec) [ -35101.3845] [ 6409] [depth= 10] . ( 30 sec) [ -35025.8677] [ 6393] [depth= 22] . ( 31 sec) [ -34976.5487] [ 6372] [depth= 15] . ( 31 sec) [ -34926.4429] [ 6352] [depth= 15] . ( 31 sec) [ -34902.3489] [ 6349] [depth= 3] . ( 32 sec) [ -34902.0558] [ 6349] [depth= 1] . ( 32 sec) [ -34820.0088] [ 6350] [depth= 11] . ( 32 sec) [ -34733.2144] [ 6328] [depth= 20] . ( 32 sec) [ -34660.1386] [ 6309] [depth= 16] . ( 33 sec) [ -34060.8360] [ 6284] [depth= 15] . ( 33 sec) [ -33980.2840] [ 6292] [depth= 1] . ( 33 sec) [ -33825.7388] [ 6258] [depth= 16] . ( 34 sec) [ -33482.6074] [ 6191] [depth= 20] . ( 34 sec) [ -33008.9987] [ 6196] [depth= 18] . ( 34 sec) [ -32926.1446] [ 6185] [depth= 19] . ( 34 sec) [ -32889.2080] [ 6173] [depth= 16] . ( 35 sec) [ -32863.6498] [ 6169] [depth= 7] . ( 35 sec) [ -32860.8947] [ 6170] [depth= 2] . ( 35 sec) [ -32746.4389] [ 6166] [depth= 9] . ( 35 sec) [ -32645.0345] [ 6157] [depth= 6] . ( 36 sec) [ -32436.4059] [ 6105] [depth= 16] . ( 36 sec) [ -32432.6386] [ 6104] [depth= 1] . ( 36 sec) [ -32119.2921] [ 6105] [depth= 23] . ( 37 sec) [ -32100.9253] [ 6090] [depth= 3] . ( 38 sec) [ -32055.9132] [ 6067] [depth= 8] . ( 38 sec) [ -32037.9436] [ 6079] [depth= 2] . ( 38 sec) [ -32024.8183] [ 6072] [depth= 14] . ( 38 sec) [ -32007.9326] [ 6058] [depth= 1] . ( 39 sec) [ -31774.4060] [ 6000] [depth= 18] . ( 39 sec) [ -31736.3957] [ 5993] [depth= 1] . ( 39 sec) [ -31723.0671] [ 5992] [depth= 1] . ( 39 sec) [ -31683.3186] [ 5982] [depth= 2] . ( 40 sec) [ -31682.9800] [ 5982] [depth= 1] . ( 40 sec) [ -31658.8278] [ 5979] [depth= 7] . ( 40 sec) [ -31653.8395] [ 5980] [depth= 1] . ( 40 sec) [ -31645.0292] [ 5976] [depth= 2] . ( 41 sec) [ -31600.6473] [ 5964] [depth= 14] . ( 41 sec) [ -31561.6531] [ 5966] [depth= 1] . ( 41 sec) [ -31533.7259] [ 5962] [depth= 4] . ( 42 sec) [ -31389.4314] [ 5912] [depth= 2] . ( 42 sec) [ -31387.2151] [ 5914] [depth= 1] . ( 42 sec) [ -31310.7892] [ 5900] [depth= 4] . ( 42 sec) [ -31310.3926] [ 5900] [depth= 1] . ( 43 sec) [ -31308.3363] [ 5899] [depth= 2] . ( 43 sec) [ -31294.4287] [ 5899] [depth= 8] . ( 43 sec) [ -31292.3460] [ 5899] [depth= 1] . ( 43 sec) [ -31174.8106] [ 5911] [depth= 9] . ( 44 sec) [ -31099.7947] [ 5897] [depth= 8] . ( 44 sec) [ -31003.7960] [ 5855] [depth= 19] . ( 44 sec) [ -30993.0289] [ 5858] [depth= 8] . ( 45 sec) [ -30976.6501] [ 5866] [depth= 19] . ( 45 sec) [ -30967.1895] [ 5861] [depth= 6] . ( 45 sec) [ -30965.5415] [ 5859] [depth= 1] . ( 45 sec) [ -30925.7734] [ 5846] [depth= 5] . ( 46 sec) [ -30922.4963] [ 5842] [depth= 2] . ( 46 sec) [ -30912.3734] [ 5837] [depth= 1] . ( 46 sec) [ -30883.8080] [ 5833] [depth= 9] . ( 46 sec) [ -30877.5594] [ 5803] [depth= 17] . ( 46 sec) [ -30875.7352] [ 5795] [depth= 1] . ( 47 sec) [ -30795.4634] [ 5771] [depth= 3] . ( 47 sec) [ -30680.4552] [ 5744] [depth= 2] . ( 47 sec) [ -30636.3843] [ 5705] [depth= 12] . ( 48 sec) [ -29848.0567] [ 5660] [depth= 12] . ( 48 sec) [ -29726.9625] [ 5664] [depth= 16] . ( 48 sec) [ -29401.2795] [ 5660] [depth= 2] . ( 49 sec) [ -29393.4590] [ 5659] [depth= 1] . ( 49 sec) [ -29377.9555] [ 5655] [depth= 2] . ( 49 sec) [ -29333.7111] [ 5644] [depth= 3] . ( 49 sec) [ -27461.7553] [ 5597] [depth= 11] . ( 50 sec) [ -27129.8249] [ 5536] [depth= 17] . ( 50 sec) [ -26746.3301] [ 5446] [depth= 11] . ( 51 sec) [ -26738.2489] [ 5444] [depth= 5] . ( 51 sec) [ -26734.8989] [ 5444] [depth= 1] . ( 51 sec) [ -26725.8306] [ 5441] [depth= 3] . ( 51 sec) [ -26672.1753] [ 5445] [depth= 1] . ( 51 sec) [ -26583.0327] [ 5427] [depth= 9] . ( 52 sec) [ -26530.5420] [ 5410] [depth= 4] . ( 52 sec) [ -26524.5362] [ 5400] [depth= 7] . ( 52 sec) [ -26481.4352] [ 5395] [depth= 12] . ( 53 sec) [ -26472.0873] [ 5395] [depth= 17] . ( 53 sec) [ -26449.8752] [ 5391] [depth= 14] . ( 53 sec) [ -26384.4658] [ 5381] [depth= 13] . ( 54 sec) [ -26322.2081] [ 5372] [depth= 1] . ( 54 sec) [ -26301.2813] [ 5354] [depth= 7] . ( 54 sec) [ -26286.1363] [ 5353] [depth= 17] . ( 55 sec) [ -26280.0756] [ 5355] [depth= 1] . ( 55 sec) [ -26278.8862] [ 5357] [depth= 1] . ( 55 sec) [ -26261.0031] [ 5356] [depth= 1] . ( 56 sec) [ -26260.5815] [ 5354] [depth= 2] . ( 56 sec) [ -26251.6537] [ 5353] [depth= 1] . ( 56 sec) [ -26231.5746] [ 5348] [depth= 17] . ( 57 sec) [ -26137.9790] [ 5315] [depth= 17] . ( 57 sec) [ -26137.7476] [ 5316] [depth= 1] . ( 58 sec) [ -26132.7014] [ 5314] [depth= 2] . ( 58 sec) [ -26123.0977] [ 5314] [depth= 1] . ( 58 sec) [ -26117.8800] [ 5312] [depth= 1] . ( 58 sec) [ -26099.7979] [ 5310] [depth= 3] . ( 59 sec) [ -26092.3921] [ 5309] [depth= 4] . ( 59 sec) [ -26078.2057] [ 5305] [depth= 4] . ( 59 sec) [ -26043.7383] [ 5294] [depth= 4] . ( 59 sec) [ -26014.7709] [ 5289] [depth= 13] . ( 60 sec) [ -26011.3484] [ 5286] [depth= 8] . ( 60 sec) [ -25983.2190] [ 5289] [depth= 2] . ( 60 sec) [ -25978.2612] [ 5289] [depth= 1] . ( 61 sec) [ -25333.0332] [ 5274] [depth= 11] . ( 61 sec) [ -25027.4427] [ 5134] [depth= 15] . ( 61 sec) [ -25018.5737] [ 5137] [depth= 1] . ( 61 sec) [ -24880.5664] [ 5111] [depth= 14] . ( 61 sec) [ -24849.0957] [ 5095] [depth= 15] . ( 62 sec) [ -24832.2714] [ 5085] [depth= 14] . ( 62 sec) [ -24824.7057] [ 5086] [depth= 1] . ( 62 sec) [ -24647.9572] [ 5031] [depth= 15] . ( 63 sec) [ -24636.7126] [ 5028] [depth= 15] . ( 63 sec) [ -24592.6682] [ 5028] [depth= 1] . ( 64 sec) [ -24419.6047] [ 4987] [depth= 2] . ( 64 sec) [ -24392.9742] [ 4985] [depth= 12] . ( 64 sec) [ -24384.3405] [ 4978] [depth= 1] . ( 64 sec) [ -24325.7711] [ 4957] [depth= 12] . ( 65 sec) [ -23922.0234] [ 4946] [depth= 1] . ( 65 sec) [ -23911.0487] [ 4940] [depth= 1] . ( 65 sec) [ -23910.2265] [ 4940] [depth= 1] . ( 65 sec) [ -23884.1481] [ 4928] [depth= 1] . ( 66 sec) [ -23883.3339] [ 4928] [depth= 11] . ( 66 sec) [ -23852.9294] [ 4929] [depth= 1] . ( 66 sec) [ -23813.0505] [ 4923] [depth= 8] . ( 67 sec) [ -23779.3709] [ 4916] [depth= 15] . ( 67 sec) [ -23774.1878] [ 4913] [depth= 1] . ( 67 sec) [ -23765.9451] [ 4912] [depth= 12] . ( 67 sec) [ -23763.4982] [ 4911] [depth= 1] . ( 68 sec) [ -23728.6156] [ 4899] [depth= 2] . ( 68 sec) [ -23470.4212] [ 4897] [depth= 1] . ( 68 sec) [ -23441.1920] [ 4885] [depth= 17] . ( 68 sec) [ -23428.3955] [ 4883] [depth= 1] . ( 69 sec) [ -23368.4946] [ 4854] [depth= 10] . ( 69 sec) [ -23368.4597] [ 4854] [depth= 1] . ( 69 sec) [ -23365.0505] [ 4850] [depth= 3] . ( 70 sec) [ -23360.2398] [ 4847] [depth= 2] . ( 70 sec) [ -23356.3303] [ 4846] [depth= 1] . ( 71 sec) [ -23333.0035] [ 4835] [depth= 6] . ( 71 sec) [ -23316.0994] [ 4835] [depth= 1] . ( 71 sec) [ -23310.9589] [ 4828] [depth= 18] . ( 71 sec) [ -23289.2516] [ 4779] [depth= 2] . ( 72 sec) [ -23286.8817] [ 4780] [depth= 1] . ( 72 sec) [ -23266.8755] [ 4782] [depth= 1] . ( 72 sec) [ -23139.4437] [ 4716] [depth= 1] . ( 73 sec) [ -23123.1394] [ 4708] [depth= 17] . ( 74 sec) [ -23105.3519] [ 4723] [depth= 1] . ( 74 sec) [ -23094.3148] [ 4718] [depth= 2] . ( 74 sec) [ -22996.2091] [ 4690] [depth= 4] . ( 75 sec) [ -22990.0006] [ 4686] [depth= 3] . ( 75 sec) [ -22958.5595] [ 4687] [depth= 4] . ( 75 sec) [ -22958.1610] [ 4689] [depth= 1] . ( 75 sec) [ -22948.7072] [ 4684] [depth= 18] . ( 76 sec) [ -22936.5500] [ 4681] [depth= 2] . ( 76 sec) [ -22920.8500] [ 4677] [depth= 16] . ( 76 sec) [ -22660.4831] [ 4567] [depth= 3] . ( 77 sec) [ -22653.7656] [ 4568] [depth= 2] . ( 77 sec) [ -22646.5580] [ 4563] [depth= 1] . ( 77 sec) [ -22641.6176] [ 4561] [depth= 1] . ( 78 sec) [ -22499.5873] [ 4523] [depth= 10] . ( 78 sec) [ -22439.5345] [ 4514] [depth= 5] . ( 78 sec) [ -22388.7739] [ 4514] [depth= 1] . ( 79 sec) [ -22388.0420] [ 4517] [depth= 1] . ( 80 sec) [ -22368.1784] [ 4517] [depth= 3] . ( 80 sec) [ -22359.1632] [ 4514] [depth= 8] . ( 80 sec) [ -22356.4790] [ 4514] [depth= 1] . ( 81 sec) [ -22355.1164] [ 4513] [depth= 2] . ( 82 sec) [ -22354.5897] [ 4513] [depth= 1] . ( 82 sec) [ -22137.5742] [ 4414] [depth= 10] . ( 82 sec) [ -22114.6511] [ 4409] [depth= 11] . ( 83 sec) [ -22112.4516] [ 4409] [depth= 1] . ( 83 sec) [ -22089.2046] [ 4400] [depth= 11] . ( 84 sec) [ -22087.4618] [ 4400] [depth= 2] . ( 84 sec) [ -22087.2734] [ 4401] [depth= 1] . ( 84 sec) [ -22086.1856] [ 4399] [depth= 10] . ( 84 sec) [ -22072.4969] [ 4393] [depth= 9] . ( 85 sec) [ -22061.0808] [ 4394] [depth= 8] . ( 85 sec) [ -22060.7719] [ 4394] [depth= 1] . ( 86 sec) [ -22057.7908] [ 4398] [depth= 19] . ( 86 sec) [ -22036.8355] [ 4395] [depth= 1] . ( 87 sec) [ -22024.6228] [ 4395] [depth= 19] . ( 87 sec) [ -22014.1717] [ 4396] [depth= 2] . ( 87 sec) [ -21991.6817] [ 4397] [depth= 1] . ( 87 sec) [ -21988.6593] [ 4395] [depth= 1] . ( 88 sec) [ -21949.4660] [ 4369] [depth= 18] . ( 88 sec) [ -21949.1963] [ 4369] [depth= 1] . ( 88 sec) [ -21947.4463] [ 4369] [depth= 4] . ( 89 sec) [ -21937.9028] [ 4368] [depth= 8] . ( 89 sec) [ -21936.7204] [ 4368] [depth= 3] . ( 89 sec) [ -21933.4080] [ 4369] [depth= 1] . ( 89 sec) [ -21922.8849] [ 4365] [depth= 2] . ( 91 sec) [ -21921.2441] [ 4365] [depth= 1] . ( 92 sec) [ -21915.3725] [ 4363] [depth= 10] . ( 92 sec) [ -21907.1719] [ 4362] [depth= 18] . ( 92 sec) [ -21903.7906] [ 4361] [depth= 1] . ( 93 sec) [ -21884.7725] [ 4358] [depth= 4] . ( 93 sec) [ -21883.1956] [ 4359] [depth= 1] . ( 94 sec) [ -21878.5680] [ 4357] [depth= 1] . ( 94 sec) [ -21870.1676] [ 4355] [depth= 5] . ( 95 sec) [ -21868.4228] [ 4358] [depth= 2] . ( 96 sec) [ -21841.7215] [ 4354] [depth= 9] . ( 96 sec) [ -21790.7690] [ 4312] [depth= 4] . ( 96 sec) [ -21786.5905] [ 4309] [depth= 12] . ( 97 sec) [ -21781.4628] [ 4308] [depth= 11] . ( 97 sec) [ -21775.2329] [ 4307] [depth= 1] . ( 98 sec) [ -21763.0740] [ 4299] [depth= 11] . ( 98 sec) [ -21756.1514] [ 4300] [depth= 1] . ( 98 sec) [ -21753.1127] [ 4298] [depth= 1] . ( 99 sec) [ -21750.6245] [ 4297] [depth= 2] . ( 99 sec) [ -21735.0504] [ 4293] [depth= 3] . ( 100 sec) [ -21733.9961] [ 4291] [depth= 2] . ( 100 sec) [ -21729.2023] [ 4288] [depth= 3] . ( 102 sec) [ -21725.8755] [ 4289] [depth= 1] . ( 102 sec) [ -21629.1592] [ 4266] [depth= 2] . ( 102 sec) [ -21599.7909] [ 4249] [depth= 10] . ( 103 sec) [ -21541.7504] [ 4225] [depth= 9] . ( 105 sec) [ -21160.4574] [Branch lengths ] . ( 111 sec) [ -21133.3917] [GTR parameters ] . ( 112 sec) [ -21076.4570] [Alpha ][ 0.344567] . ( 112 sec) [ -21067.3183] [ 4222] [depth= 12] . ( 112 sec) [ -21048.7190] [ 4218] [depth= 11] . ( 113 sec) [ -21046.0512] [ 4215] [depth= 3] . ( 113 sec) [ -21030.0457] [ 4212] [depth= 3] . ( 115 sec) [ -21026.4837] [ 4210] [depth= 1] . ( 117 sec) [ -21026.2071] [ 4212] [depth= 1] . ( 118 sec) [ -21025.1867] [ 4210] [depth= 1] . ( 119 sec) [ -21025.0683] [ 4210] [depth= 1] . ( 120 sec) [ -21013.0570] [ 4205] [depth= 8] . ( 121 sec) [ -21002.8990] [ 4200] [depth= 2] . ( 121 sec) [ -20991.7537] [ 4195] [depth= 4] . ( 122 sec) [ -20967.2963] [ 4175] [depth= 13] . ( 122 sec) [ -20964.3850] [ 4173] [depth= 1] . ( 123 sec) [ -20962.5602] [ 4172] [depth= 1] . ( 123 sec) [ -20961.5957] [ 4173] [depth= 1] . ( 124 sec) [ -20959.3903] [ 4173] [depth= 1] . ( 125 sec) [ -20942.6411] [ 4163] [depth= 3] . ( 125 sec) [ -20912.8804] [ 4153] [depth= 12] . ( 125 sec) [ -20912.1420] [ 4153] [depth= 2] . ( 126 sec) [ -20911.0310] [ 4155] [depth= 1] . ( 127 sec) [ -20909.4534] [ 4154] [depth= 1] . ( 128 sec) [ -20884.5911] [ 4146] [depth= 3] . ( 128 sec) [ -20880.9859] [ 4142] [depth= 1] . ( 128 sec) [ -20878.7018] [ 4142] [depth= 1] . ( 129 sec) [ -20878.6552] [ 4145] [depth= 6] . ( 129 sec) [ -20877.2449] [ 4145] [depth= 1] . ( 130 sec) [ -20874.2203] [ 4148] [depth= 1] . ( 131 sec) [ -20857.1871] [ 4140] [depth= 17] . ( 132 sec) [ -20843.5531] [ 4133] [depth= 5] . ( 132 sec) [ -20836.1710] [ 4128] [depth= 3] . ( 132 sec) [ -20824.5146] [ 4125] [depth= 11] . ( 133 sec) [ -20816.2149] [ 4129] [depth= 1] . ( 134 sec) [ -20813.8062] [ 4127] [depth= 1] . ( 135 sec) [ -20813.0849] [ 4125] [depth= 3] . ( 138 sec) [ -20802.0381] [ 4120] [depth= 6] . ( 139 sec) [ -20795.6736] [ 4117] [depth= 1] . ( 139 sec) [ -20793.7450] [ 4116] [depth= 1] . ( 140 sec) [ -20793.6412] [ 4115] [depth= 4] . ( 140 sec) [ -20779.0973] [ 4113] [depth= 4] . ( 140 sec) [ -20768.9001] [ 4115] [depth= 1] . ( 144 sec) [ -20746.1063] [ 4099] [depth= 9] . ( 145 sec) [ -20745.4158] [ 4094] [depth= 1] . ( 145 sec) [ -20735.9124] [ 4092] [depth= 2] . ( 148 sec) [ -20735.0174] [ 4092] [depth= 1] . ( 148 sec) [ -20729.6736] [ 4090] [depth= 2] . ( 149 sec) [ -20728.4463] [ 4091] [depth= 1] . ( 149 sec) [ -20726.0472] [ 4091] [depth= 2] . ( 149 sec) [ -20725.7384] [ 4091] [depth= 6] . ( 150 sec) [ -20724.6798] [ 4095] [depth= 1] . ( 151 sec) [ -20721.8694] [ 4094] [depth= 13] . ( 152 sec) [ -20698.8501] [ 4094] [depth= 20] . ( 153 sec) [ -20688.6647] [ 4084] [depth= 1] . ( 154 sec) [ -20686.7184] [ 4083] [depth= 1] . ( 155 sec) [ -20678.4241] [ 4082] [depth= 1] . ( 155 sec) [ -20677.0948] [ 4082] [depth= 1] . ( 155 sec) [ -20673.6245] [ 4082] [depth= 1] . ( 156 sec) [ -20658.0124] [ 4074] [depth= 8] . ( 156 sec) [ -20647.5877] [ 4070] [depth= 7] . ( 156 sec) [ -20634.0367] [ 4063] [depth= 7] . ( 157 sec) [ -20633.0128] [ 4063] [depth= 1] . ( 157 sec) [ -20632.7976] [ 4063] [depth= 2] . ( 162 sec) [ -20632.0170] [ 4066] [depth= 1] . ( 164 sec) [ -20630.1950] [ 4068] [depth= 1] . ( 167 sec) [ -20587.4244] [ 4064] [depth= 15] . ( 167 sec) [ -20587.0838] [ 4064] [depth= 1] . ( 168 sec) [ -20571.2464] [ 4069] [depth= 1] . ( 168 sec) [ -20566.8070] [ 4066] [depth= 3] . ( 174 sec) [ -20565.7468] [ 4067] [depth= 1] . ( 175 sec) [ -20565.2037] [ 4067] [depth= 1] . ( 175 sec) [ -20564.5168] [ 4065] [depth= 3] . ( 176 sec) [ -20564.1203] [ 4070] [depth= 1] . ( 176 sec) [ -20561.8757] [ 4073] [depth= 7] . ( 177 sec) [ -20561.2371] [ 4073] [depth= 1] . ( 177 sec) [ -20560.7328] [ 4072] [depth= 1] . ( 178 sec) [ -20560.7177] [ 4072] [depth= 1] . ( 178 sec) [ -20560.4482] [ 4073] [depth= 1] . ( 179 sec) [ -20553.5330] [ 4071] [depth= 7] . ( 180 sec) [ -20552.8809] [ 4074] [depth= 1] . ( 181 sec) [ -20552.3126] [ 4076] [depth= 1] . ( 181 sec) [ -20552.2937] [ 4075] [depth= 1] . ( 183 sec) [ -20552.2433] [ 4075] [depth= 1] . ( 184 sec) [ -20542.7104] [ 4073] [depth= 17] . ( 184 sec) [ -20539.2443] [ 4068] [depth= 1] . ( 185 sec) [ -20535.9839] [ 4068] [depth= 4] . ( 185 sec) [ -20528.9174] [ 4062] [depth= 6] . ( 185 sec) [ -20525.4972] [ 4061] [depth= 1] . ( 187 sec) [ -20525.0688] [ 4061] [depth= 1] . ( 188 sec) [ -20524.0195] [ 4063] [depth= 1] . ( 188 sec) [ -20523.8717] [ 4064] [depth= 3] . ( 192 sec) [ -20523.8077] [ 4064] [depth= 1] . ( 192 sec) [ -20522.5850] [ 4065] [depth= 1] . ( 193 sec) [ -20519.6881] [ 4064] [depth= 1] . ( 193 sec) [ -20513.9550] [ 4062] [depth= 4] . ( 194 sec) [ -20513.5810] [ 4062] [depth= 1] . ( 195 sec) [ -20511.8944] [ 4057] [depth= 5] . ( 200 sec) [ -20508.5095] [ 4056] [depth= 2] . ( 202 sec) [ -20508.1452] [ 4056] [depth= 1] . ( 205 sec) [ -20506.8407] [ 4055] [depth= 1] . ( 211 sec) [ -20501.4219] [ 4054] [depth= 5] . ( 213 sec) [ -20499.6631] [ 4055] [depth= 1] . ( 213 sec) [ -20496.0795] [ 4059] [depth= 1] . ( 214 sec) [ -20494.1538] [ 4058] [depth= 1] . ( 216 sec) [ -20441.5347] [Branch lengths ] . ( 220 sec) [ -20441.2016] [GTR parameters ] . ( 221 sec) [ -20441.1796] [Alpha ][ 0.342307] . ( 228 sec) [ -20440.9024] [ 4058] [depth= 1] . ( 230 sec) [ -20440.8894] [ 4058] [depth= 1] . ( 243 sec) [ -20438.9133] [ 4059] [depth= 2] . ( 250 sec) [ -20438.3146] [ 4058] [depth= 1] . ( 251 sec) [ -20437.8596] [ 4059] [depth= 2] . ( 252 sec) [ -20436.9609] [ 4060] [depth= 1] . ( 256 sec) [ -20436.3716] [ 4061] [depth= 1] . ( 261 sec) [ -20436.0348] [ 4062] [depth= 2] . ( 261 sec) [ -20434.8350] [ 4054] [depth= 10] . ( 272 sec) [ -20434.4310] [ 4054] [depth= 1] . ( 272 sec) [ -20433.8282] [ 4052] [depth= 1] . ( 275 sec) [ -20433.4053] [ 4053] [depth= 2] . ( 283 sec) [ -20430.8494] [ 4051] [depth= 1] . ( 290 sec) [ -20430.3540] [ 4051] [depth= 1] . ( 291 sec) [ -20428.7937] [ 4052] [depth= 3] . ( 297 sec) [ -20428.7812] [ 4052] [depth= 1] . ( 308 sec) [ -20428.7274] [ 4052] [depth= 1] . ( 309 sec) [ -20424.2582] [ 4055] [depth= 1] . ( 310 sec) [ -20423.8243] [ 4055] [depth= 1] . ( 319 sec) [ -20421.5265] [ 4052] [depth= 1] . ( 324 sec) [ -20420.3468] [ 4051] [depth= 1] . ( 325 sec) [ -20419.5289] [ 4051] [depth= 1] . ( 326 sec) [ -20419.4037] [ 4052] [depth= 1] . ( 337 sec) [ -20419.3756] [ 4049] [depth= 1] . ( 338 sec) [ -20419.0418] [ 4049] [depth= 1] . ( 349 sec) [ -20410.0764] [Branch lengths ] . ( 352 sec) [ -20410.0117] [GTR parameters ] . ( 352 sec) [ -20410.0092] [Alpha ][ 0.341381] . ( 371 sec) [ -20409.7236] [ 4052] [depth= 1] . ( 465 sec) [ -20409.6823] [ 4051] [depth= 1] . ( 488 sec) [ -20408.3735] [Branch lengths ] . ( 489 sec) [ -20408.3732] [GTR parameters ] . ( 490 sec) [ -20408.3732] [Alpha ][ 0.341381] . ( 626 sec) [ -20407.2677] [Branch lengths ] . ( 878 sec) [ -20407.1197] [Branch lengths ] . ( 878 sec) [ -20407.1197] [Topology ] . ( 883 sec) [ -20407.0071] [Topology ] . ( 888 sec) [ -20406.9205] [Topology ] . ( 893 sec) [ -20406.9023] [Topology ] . ( 898 sec) [ -20406.8973] [Topology ] . ( 904 sec) [ -20406.8958] [Topology ] . ( 909 sec) [ -20406.8953] [Topology ] . ( 912 sec) [ -20406.8889] [GTR parameters ] . ( 913 sec) [ -20406.8880] [Alpha ][ 0.341785] . ( 913 sec) [ -20406.8880] [Topology ] . ( 918 sec) [ -20406.8875] [Topology ] . ( 919 sec) [ -20406.8875] [GTR parameters ] . ( 919 sec) [ -20406.8875] [Alpha ][ 0.341785] . Checking for NNIs, optimizing five branches... . ( 956 sec) [ -20406.3623] [Topology ] . Checking for NNIs, optimizing five branches... . ( 992 sec) [ -20406.1300] [Topology ] . Checking for NNIs, optimizing five branches... . ( 1029 sec) [ -20405.0487] [Topology ] . Checking for NNIs, optimizing five branches... . Log likelihood of the current tree: -20405.048694. . [Random start 2/ 5] . Computing pairwise distances... . Building BioNJ tree... . Randomising the tree... . Maximizing likelihood (using SPR moves)... . ( 6 sec) [ -42484.0193] [GTR parameters ] . ( 6 sec) [ -41489.0846] [Alpha ][ 0.500960] . ( 7 sec) [ -41367.4981] [ 7418] [depth= 10] . ( 7 sec) [ -41293.3660] [ 7399] [depth= 17] . ( 7 sec) [ -41273.2051] [ 7393] [depth= 5] . ( 8 sec) [ -41191.8432] [ 7368] [depth= 1] . ( 8 sec) [ -41007.0805] [ 7361] [depth= 7] . ( 8 sec) [ -40889.2599] [ 7359] [depth= 6] . ( 8 sec) [ -40871.5690] [ 7357] [depth= 6] . ( 9 sec) [ -40804.5348] [ 7337] [depth= 1] . ( 9 sec) [ -40753.4634] [ 7334] [depth= 5] . ( 9 sec) [ -40735.5026] [ 7331] [depth= 12] . ( 9 sec) [ -40704.9229] [ 7328] [depth= 5] . ( 10 sec) [ -40651.8498] [ 7320] [depth= 19] . ( 10 sec) [ -40619.8427] [ 7323] [depth= 2] . ( 10 sec) [ -40536.6464] [ 7306] [depth= 3] . ( 10 sec) [ -40287.0476] [ 7297] [depth= 13] . ( 10 sec) [ -40228.5914] [ 7291] [depth= 2] . ( 11 sec) [ -40165.8774] [ 7272] [depth= 2] . ( 11 sec) [ -40129.5258] [ 7261] [depth= 15] . ( 11 sec) [ -40088.0622] [ 7254] [depth= 23] . ( 11 sec) [ -39794.3628] [ 7260] [depth= 1] . ( 12 sec) [ -39719.4877] [ 7245] [depth= 2] . ( 12 sec) [ -39586.4963] [ 7232] [depth= 26] . ( 12 sec) [ -39203.3009] [ 7155] [depth= 14] . ( 13 sec) [ -38724.3212] [ 7139] [depth= 22] . ( 13 sec) [ -38712.0879] [ 7136] [depth= 1] . ( 13 sec) [ -38516.4395] [ 7138] [depth= 1] . ( 14 sec) [ -38006.8248] [ 6977] [depth= 13] . ( 14 sec) [ -37980.9153] [ 6971] [depth= 19] . ( 14 sec) [ -37922.4506] [ 6958] [depth= 7] . ( 14 sec) [ -37891.7034] [ 6958] [depth= 2] . ( 15 sec) [ -37840.3630] [ 6947] [depth= 4] . ( 15 sec) [ -37334.9150] [ 6935] [depth= 8] . ( 15 sec) [ -37318.6732] [ 6933] [depth= 13] . ( 15 sec) [ -37293.5656] [ 6930] [depth= 8] . ( 15 sec) [ -37281.3529] [ 6928] [depth= 6] . ( 16 sec) [ -37182.3521] [ 6933] [depth= 1] . ( 16 sec) [ -37153.4342] [ 6927] [depth= 16] . ( 16 sec) [ -37097.1426] [ 6918] [depth= 2] . ( 17 sec) [ -37043.5525] [ 6920] [depth= 1] . ( 17 sec) [ -36910.9266] [ 6897] [depth= 14] . ( 17 sec) [ -36024.1235] [ 6889] [depth= 3] . ( 17 sec) [ -35963.9724] [ 6886] [depth= 7] . ( 18 sec) [ -35835.3703] [ 6879] [depth= 2] . ( 18 sec) [ -35772.4358] [ 6863] [depth= 1] . ( 18 sec) [ -35647.6968] [ 6824] [depth= 10] . ( 18 sec) [ -35634.7566] [ 6822] [depth= 10] . ( 19 sec) [ -35620.9645] [ 6820] [depth= 3] . ( 19 sec) [ -35597.2421] [ 6818] [depth= 2] . ( 19 sec) [ -35520.9523] [ 6803] [depth= 14] . ( 19 sec) [ -35467.0906] [ 6794] [depth= 11] . ( 20 sec) [ -35443.3130] [ 6791] [depth= 11] . ( 20 sec) [ -35385.6234] [ 6753] [depth= 11] . ( 20 sec) [ -35060.7760] [ 6716] [depth= 10] . ( 21 sec) [ -35007.0201] [ 6704] [depth= 11] . ( 21 sec) [ -34944.8299] [ 6701] [depth= 17] . ( 21 sec) [ -34805.9533] [ 6721] [depth= 1] . ( 21 sec) [ -34524.4201] [ 6675] [depth= 16] . ( 22 sec) [ -34514.8578] [ 6676] [depth= 1] . ( 22 sec) [ -33998.4989] [ 6652] [depth= 6] . ( 22 sec) [ -33694.2885] [ 6611] [depth= 9] . ( 22 sec) [ -33551.2872] [ 6556] [depth= 14] . ( 23 sec) [ -33117.3391] [ 6453] [depth= 6] . ( 23 sec) [ -32997.8606] [ 6445] [depth= 7] . ( 23 sec) [ -32997.1790] [ 6442] [depth= 11] . ( 23 sec) [ -32975.6859] [ 6441] [depth= 4] . ( 24 sec) [ -32935.4836] [ 6440] [depth= 2] . ( 24 sec) [ -32929.3086] [ 6441] [depth= 1] . ( 24 sec) [ -32926.5895] [ 6439] [depth= 1] . ( 25 sec) [ -32848.9328] [ 6423] [depth= 21] . ( 25 sec) [ -32738.9885] [ 6399] [depth= 25] . ( 25 sec) [ -32683.2345] [ 6381] [depth= 2] . ( 25 sec) [ -32439.2063] [ 6381] [depth= 13] . ( 26 sec) [ -32438.9612] [ 6376] [depth= 10] . ( 26 sec) [ -32379.4814] [ 6371] [depth= 4] . ( 26 sec) [ -32315.5689] [ 6364] [depth= 8] . ( 27 sec) [ -32293.7983] [ 6363] [depth= 1] . ( 27 sec) [ -32263.7438] [ 6353] [depth= 1] . ( 27 sec) [ -32255.1157] [ 6352] [depth= 2] . ( 27 sec) [ -32177.7620] [ 6349] [depth= 14] . ( 28 sec) [ -32110.9272] [ 6332] [depth= 2] . ( 28 sec) [ -32074.7640] [ 6322] [depth= 11] . ( 28 sec) [ -32041.3296] [ 6317] [depth= 8] . ( 29 sec) [ -32036.2003] [ 6316] [depth= 8] . ( 29 sec) [ -31639.6776] [ 6312] [depth= 10] . ( 29 sec) [ -31605.8129] [ 6310] [depth= 1] . ( 30 sec) [ -31579.7067] [ 6307] [depth= 4] . ( 30 sec) [ -31573.1291] [ 6308] [depth= 3] . ( 30 sec) [ -31480.8146] [ 6305] [depth= 7] . ( 31 sec) [ -31406.1687] [ 6266] [depth= 12] . ( 31 sec) [ -30882.3033] [ 6241] [depth= 16] . ( 31 sec) [ -30713.2822] [ 6225] [depth= 10] . ( 32 sec) [ -30671.6348] [ 6217] [depth= 19] . ( 32 sec) [ -30488.7629] [ 6187] [depth= 18] . ( 32 sec) [ -30473.2067] [ 6188] [depth= 19] . ( 32 sec) [ -30467.0517] [ 6184] [depth= 2] . ( 33 sec) [ -30462.4609] [ 6184] [depth= 1] . ( 33 sec) [ -30454.0422] [ 6173] [depth= 8] . ( 33 sec) [ -30336.8017] [ 6171] [depth= 2] . ( 33 sec) [ -30257.2340] [ 6167] [depth= 13] . ( 34 sec) [ -30208.0670] [ 6186] [depth= 1] . ( 34 sec) [ -30129.0578] [ 6154] [depth= 18] . ( 34 sec) [ -29989.4469] [ 6132] [depth= 15] . ( 35 sec) [ -29921.4872] [ 6111] [depth= 7] . ( 35 sec) [ -29862.6228] [ 6106] [depth= 16] . ( 35 sec) [ -29764.0123] [ 6091] [depth= 5] . ( 36 sec) [ -29694.9245] [ 6094] [depth= 1] . ( 36 sec) [ -29571.0983] [ 6085] [depth= 3] . ( 36 sec) [ -29560.2695] [ 6099] [depth= 1] . ( 36 sec) [ -29324.8884] [ 5984] [depth= 8] . ( 37 sec) [ -29182.7203] [ 5972] [depth= 20] . ( 37 sec) [ -29119.4897] [ 5964] [depth= 1] . ( 37 sec) [ -29119.1667] [ 5965] [depth= 1] . ( 37 sec) [ -29088.3516] [ 5963] [depth= 3] . ( 38 sec) [ -29053.3293] [ 5956] [depth= 7] . ( 38 sec) [ -28979.6776] [ 5914] [depth= 16] . ( 38 sec) [ -28977.7685] [ 5914] [depth= 1] . ( 39 sec) [ -28976.2614] [ 5912] [depth= 8] . ( 39 sec) [ -28957.6353] [ 5911] [depth= 8] . ( 39 sec) [ -28757.7301] [ 5888] [depth= 18] . ( 39 sec) [ -28691.8681] [ 5891] [depth= 1] . ( 40 sec) [ -28666.1325] [ 5885] [depth= 4] . ( 40 sec) [ -28522.4456] [ 5861] [depth= 15] . ( 40 sec) [ -28500.5749] [ 5858] [depth= 6] . ( 41 sec) [ -28487.7982] [ 5853] [depth= 1] . ( 41 sec) [ -28458.6279] [ 5845] [depth= 17] . ( 41 sec) [ -27948.8584] [ 5763] [depth= 18] . ( 41 sec) [ -27872.3970] [ 5747] [depth= 13] . ( 42 sec) [ -27870.8424] [ 5749] [depth= 2] . ( 42 sec) [ -27855.0972] [ 5737] [depth= 9] . ( 42 sec) [ -27782.4783] [ 5722] [depth= 4] . ( 43 sec) [ -27769.0422] [ 5721] [depth= 1] . ( 43 sec) [ -27763.6656] [ 5705] [depth= 7] . ( 43 sec) [ -27755.6447] [ 5704] [depth= 1] . ( 44 sec) [ -27510.8308] [ 5647] [depth= 16] . ( 44 sec) [ -27324.6602] [ 5581] [depth= 15] . ( 44 sec) [ -27293.0743] [ 5560] [depth= 17] . ( 44 sec) [ -27119.2271] [ 5557] [depth= 1] . ( 45 sec) [ -27034.1613] [ 5518] [depth= 2] . ( 45 sec) [ -27028.5789] [ 5447] [depth= 20] . ( 45 sec) [ -26793.3037] [ 5450] [depth= 2] . ( 45 sec) [ -26674.6878] [ 5431] [depth= 17] . ( 46 sec) [ -26344.3024] [ 5347] [depth= 4] . ( 46 sec) [ -26270.3172] [ 5327] [depth= 4] . ( 46 sec) [ -26139.2530] [ 5311] [depth= 4] . ( 46 sec) [ -26138.2610] [ 5311] [depth= 1] . ( 47 sec) [ -26025.5235] [ 5265] [depth= 18] . ( 47 sec) [ -25919.7425] [ 5232] [depth= 1] . ( 47 sec) [ -25916.9815] [ 5229] [depth= 1] . ( 48 sec) [ -25780.1228] [ 5167] [depth= 7] . ( 48 sec) [ -25770.0015] [ 5159] [depth= 11] . ( 49 sec) [ -25564.0167] [ 5048] [depth= 8] . ( 49 sec) [ -25545.9077] [ 5043] [depth= 13] . ( 49 sec) [ -25538.2458] [ 5045] [depth= 1] . ( 49 sec) [ -25507.9022] [ 5038] [depth= 12] . ( 50 sec) [ -25507.0207] [ 5025] [depth= 12] . ( 50 sec) [ -25472.3856] [ 5025] [depth= 1] . ( 50 sec) [ -25461.4013] [ 5025] [depth= 1] . ( 50 sec) [ -25453.0770] [ 5024] [depth= 8] . ( 51 sec) [ -25452.1610] [ 5024] [depth= 1] . ( 51 sec) [ -25416.3435] [ 5002] [depth= 2] . ( 51 sec) [ -25139.9851] [ 4975] [depth= 16] . ( 52 sec) [ -25030.8759] [ 4956] [depth= 12] . ( 52 sec) [ -25009.4152] [ 4953] [depth= 5] . ( 52 sec) [ -24963.2774] [ 4950] [depth= 3] . ( 53 sec) [ -24953.5723] [ 4948] [depth= 2] . ( 53 sec) [ -24942.8206] [ 4947] [depth= 8] . ( 53 sec) [ -24937.5033] [ 4946] [depth= 1] . ( 54 sec) [ -24907.2740] [ 4937] [depth= 1] . ( 54 sec) [ -24797.9301] [ 4926] [depth= 10] . ( 54 sec) [ -24781.2477] [ 4921] [depth= 3] . ( 54 sec) [ -24719.5853] [ 4903] [depth= 15] . ( 55 sec) [ -24701.8970] [ 4891] [depth= 10] . ( 55 sec) [ -24680.2586] [ 4885] [depth= 15] . ( 55 sec) [ -24650.0677] [ 4876] [depth= 15] . ( 56 sec) [ -24632.0359] [ 4873] [depth= 1] . ( 56 sec) [ -24553.2281] [ 4857] [depth= 13] . ( 56 sec) [ -24491.8925] [ 4859] [depth= 1] . ( 56 sec) [ -24411.3603] [ 4808] [depth= 15] . ( 57 sec) [ -24328.4303] [ 4792] [depth= 15] . ( 57 sec) [ -24325.4624] [ 4792] [depth= 1] . ( 58 sec) [ -24313.2777] [ 4787] [depth= 1] . ( 58 sec) [ -24242.8461] [ 4772] [depth= 12] . ( 58 sec) [ -24239.0205] [ 4767] [depth= 2] . ( 59 sec) [ -24234.8148] [ 4766] [depth= 1] . ( 59 sec) [ -24234.2815] [ 4766] [depth= 1] . ( 59 sec) [ -24230.4155] [ 4766] [depth= 1] . ( 59 sec) [ -24218.7221] [ 4765] [depth= 2] . ( 60 sec) [ -24218.2588] [ 4764] [depth= 1] . ( 60 sec) [ -24150.9069] [ 4729] [depth= 10] . ( 61 sec) [ -24140.3591] [ 4726] [depth= 1] . ( 61 sec) [ -24099.5441] [ 4715] [depth= 11] . ( 61 sec) [ -24097.6261] [ 4714] [depth= 2] . ( 62 sec) [ -24084.5719] [ 4710] [depth= 4] . ( 62 sec) [ -24068.8321] [ 4705] [depth= 1] . ( 62 sec) [ -24053.8164] [ 4703] [depth= 6] . ( 62 sec) [ -24049.8347] [ 4703] [depth= 2] . ( 63 sec) [ -24007.7565] [ 4701] [depth= 3] . ( 63 sec) [ -24002.0649] [ 4701] [depth= 3] . ( 63 sec) [ -23997.7469] [ 4701] [depth= 2] . ( 63 sec) [ -23997.6310] [ 4701] [depth= 1] . ( 64 sec) [ -23963.1444] [ 4701] [depth= 1] . ( 64 sec) [ -23937.9491] [ 4694] [depth= 9] . ( 64 sec) [ -23937.1225] [ 4695] [depth= 12] . ( 65 sec) [ -23936.2414] [ 4697] [depth= 1] . ( 65 sec) [ -23932.3354] [ 4693] [depth= 1] . ( 65 sec) [ -23906.7745] [ 4677] [depth= 14] . ( 66 sec) [ -23901.8834] [ 4674] [depth= 13] . ( 66 sec) [ -23889.0470] [ 4670] [depth= 2] . ( 66 sec) [ -23876.8291] [ 4670] [depth= 12] . ( 67 sec) [ -23827.4579] [ 4661] [depth= 2] . ( 68 sec) [ -23825.2840] [ 4661] [depth= 2] . ( 68 sec) [ -23816.5875] [ 4660] [depth= 1] . ( 68 sec) [ -23810.8047] [ 4659] [depth= 1] . ( 69 sec) [ -23786.7733] [ 4660] [depth= 1] . ( 69 sec) [ -23778.5251] [ 4659] [depth= 1] . ( 69 sec) [ -23777.0001] [ 4658] [depth= 1] . ( 70 sec) [ -23708.0468] [ 4656] [depth= 9] . ( 70 sec) [ -23687.5299] [ 4652] [depth= 3] . ( 70 sec) [ -23685.8977] [ 4652] [depth= 1] . ( 71 sec) [ -23683.9395] [ 4654] [depth= 2] . ( 71 sec) [ -23682.2790] [ 4653] [depth= 4] . ( 71 sec) [ -23646.2945] [ 4648] [depth= 5] . ( 72 sec) [ -23645.9648] [ 4648] [depth= 1] . ( 72 sec) [ -23640.4090] [ 4648] [depth= 6] . ( 72 sec) [ -23639.8027] [ 4647] [depth= 8] . ( 73 sec) [ -23599.6781] [ 4628] [depth= 16] . ( 73 sec) [ -23535.6960] [ 4608] [depth= 1] . ( 73 sec) [ -23494.9723] [ 4593] [depth= 2] . ( 73 sec) [ -23426.7265] [ 4576] [depth= 2] . ( 74 sec) [ -23396.2012] [ 4569] [depth= 2] . ( 74 sec) [ -23385.9747] [ 4566] [depth= 9] . ( 74 sec) [ -23373.1085] [ 4565] [depth= 1] . ( 75 sec) [ -23363.1435] [ 4562] [depth= 1] . ( 75 sec) [ -23349.6336] [ 4563] [depth= 1] . ( 75 sec) [ -23320.0226] [ 4558] [depth= 8] . ( 75 sec) [ -23317.8899] [ 4557] [depth= 4] . ( 76 sec) [ -23316.3499] [ 4556] [depth= 2] . ( 76 sec) [ -23290.2747] [ 4550] [depth= 3] . ( 76 sec) [ -23249.0620] [ 4549] [depth= 7] . ( 77 sec) [ -23243.4022] [ 4542] [depth= 7] . ( 77 sec) [ -22620.0144] [ 4544] [depth= 1] . ( 77 sec) [ -22559.3551] [ 4507] [depth= 15] . ( 78 sec) [ -22558.9828] [ 4502] [depth= 6] . ( 78 sec) [ -22528.2777] [ 4488] [depth= 2] . ( 78 sec) [ -22487.8457] [ 4477] [depth= 5] . ( 79 sec) [ -22479.8557] [ 4476] [depth= 4] . ( 79 sec) [ -22466.9702] [ 4473] [depth= 2] . ( 79 sec) [ -22458.8872] [ 4478] [depth= 1] . ( 79 sec) [ -22419.7015] [ 4477] [depth= 2] . ( 80 sec) [ -22379.6389] [ 4469] [depth= 4] . ( 80 sec) [ -22378.7751] [ 4470] [depth= 2] . ( 81 sec) [ -22369.5179] [ 4467] [depth= 5] . ( 81 sec) [ -22369.1766] [ 4467] [depth= 1] . ( 82 sec) [ -22363.6683] [ 4466] [depth= 2] . ( 82 sec) [ -22353.6569] [ 4462] [depth= 6] . ( 83 sec) [ -22335.1179] [ 4454] [depth= 13] . ( 83 sec) [ -22323.8956] [ 4451] [depth= 1] . ( 84 sec) [ -22316.7563] [ 4448] [depth= 5] . ( 84 sec) [ -22284.4665] [ 4439] [depth= 10] . ( 84 sec) [ -22281.5223] [ 4443] [depth= 1] . ( 84 sec) [ -22247.6573] [ 4448] [depth= 2] . ( 85 sec) [ -22217.3630] [ 4432] [depth= 7] . ( 85 sec) [ -22210.8204] [ 4433] [depth= 1] . ( 85 sec) [ -22208.1756] [ 4432] [depth= 1] . ( 86 sec) [ -22206.4334] [ 4433] [depth= 1] . ( 86 sec) [ -22194.5999] [ 4434] [depth= 1] . ( 86 sec) [ -22168.2489] [ 4424] [depth= 1] . ( 87 sec) [ -22163.7008] [ 4422] [depth= 9] . ( 87 sec) [ -22145.0847] [ 4418] [depth= 18] . ( 87 sec) [ -22139.4062] [ 4420] [depth= 23] . ( 88 sec) [ -22134.3354] [ 4418] [depth= 3] . ( 88 sec) [ -22133.3292] [ 4418] [depth= 1] . ( 88 sec) [ -22131.9860] [ 4418] [depth= 1] . ( 88 sec) [ -22129.7849] [ 4417] [depth= 2] . ( 89 sec) [ -22128.7988] [ 4419] [depth= 2] . ( 89 sec) [ -22128.7704] [ 4419] [depth= 1] . ( 89 sec) [ -22128.4614] [ 4419] [depth= 1] . ( 89 sec) [ -22113.4419] [ 4412] [depth= 2] . ( 90 sec) [ -22112.1519] [ 4412] [depth= 1] . ( 90 sec) [ -22103.9679] [ 4412] [depth= 1] . ( 90 sec) [ -22069.5218] [ 4412] [depth= 1] . ( 90 sec) [ -22050.3786] [ 4409] [depth= 12] . ( 91 sec) [ -22039.5007] [ 4408] [depth= 1] . ( 91 sec) [ -22025.5138] [ 4400] [depth= 7] . ( 91 sec) [ -22004.1250] [ 4396] [depth= 5] . ( 92 sec) [ -21955.5194] [ 4375] [depth= 10] . ( 93 sec) [ -21952.8941] [ 4375] [depth= 1] . ( 93 sec) [ -21874.5215] [ 4357] [depth= 1] . ( 94 sec) [ -21835.3933] [ 4343] [depth= 1] . ( 94 sec) [ -21814.5439] [ 4342] [depth= 1] . ( 94 sec) [ -21774.9773] [ 4339] [depth= 1] . ( 94 sec) [ -21765.2767] [ 4337] [depth= 6] . ( 95 sec) [ -21763.4229] [ 4338] [depth= 1] . ( 95 sec) [ -21757.1471] [ 4336] [depth= 5] . ( 97 sec) [ -21756.8006] [ 4337] [depth= 1] . ( 97 sec) [ -21756.0610] [ 4337] [depth= 1] . ( 97 sec) [ -21752.8387] [ 4338] [depth= 1] . ( 98 sec) [ -21721.9110] [ 4324] [depth= 9] . ( 98 sec) [ -21721.5909] [ 4323] [depth= 7] . ( 99 sec) [ -21713.9996] [ 4313] [depth= 10] . ( 99 sec) [ -21623.9356] [ 4308] [depth= 8] . ( 99 sec) [ -21619.7593] [ 4306] [depth= 1] . ( 100 sec) [ -21617.9242] [ 4308] [depth= 1] . ( 100 sec) [ -21617.2866] [ 4306] [depth= 1] . ( 100 sec) [ -21611.2571] [ 4306] [depth= 1] . ( 101 sec) [ -21609.4684] [ 4306] [depth= 3] . ( 102 sec) [ -21609.0828] [ 4306] [depth= 1] . ( 102 sec) [ -21594.3223] [ 4302] [depth= 7] . ( 102 sec) [ -21592.7932] [ 4302] [depth= 1] . ( 104 sec) [ -21314.3003] [Branch lengths ] . ( 111 sec) [ -21290.4781] [GTR parameters ] . ( 112 sec) [ -21235.9794] [Alpha ][ 0.348413] . ( 112 sec) [ -21230.2722] [ 4302] [depth= 2] . ( 113 sec) [ -21229.2142] [ 4300] [depth= 2] . ( 113 sec) [ -21223.2384] [ 4298] [depth= 3] . ( 115 sec) [ -21218.2780] [ 4296] [depth= 3] . ( 115 sec) [ -21217.7017] [ 4296] [depth= 1] . ( 115 sec) [ -21213.0315] [ 4295] [depth= 1] . ( 116 sec) [ -21183.7486] [ 4289] [depth= 21] . ( 116 sec) [ -21162.6724] [ 4282] [depth= 1] . ( 117 sec) [ -21155.6461] [ 4281] [depth= 1] . ( 118 sec) [ -21145.4358] [ 4281] [depth= 19] . ( 118 sec) [ -21142.2905] [ 4283] [depth= 2] . ( 118 sec) [ -21138.8171] [ 4281] [depth= 5] . ( 119 sec) [ -21135.9179] [ 4281] [depth= 1] . ( 119 sec) [ -21130.5626] [ 4277] [depth= 1] . ( 119 sec) [ -21129.6398] [ 4278] [depth= 1] . ( 120 sec) [ -21125.7408] [ 4277] [depth= 1] . ( 120 sec) [ -21106.0321] [ 4275] [depth= 13] . ( 121 sec) [ -21105.1996] [ 4273] [depth= 7] . ( 123 sec) [ -21101.8174] [ 4275] [depth= 1] . ( 124 sec) [ -21097.7988] [ 4274] [depth= 1] . ( 125 sec) [ -21086.3150] [ 4261] [depth= 14] . ( 125 sec) [ -21081.4347] [ 4263] [depth= 1] . ( 125 sec) [ -21080.0635] [ 4262] [depth= 2] . ( 125 sec) [ -21078.1420] [ 4265] [depth= 1] . ( 127 sec) [ -21073.9843] [ 4263] [depth= 1] . ( 132 sec) [ -21066.8993] [ 4263] [depth= 2] . ( 134 sec) [ -21066.6796] [ 4263] [depth= 1] . ( 135 sec) [ -21065.5289] [ 4262] [depth= 1] . ( 136 sec) [ -21065.3063] [ 4262] [depth= 1] . ( 137 sec) [ -21065.0775] [ 4262] [depth= 1] . ( 138 sec) [ -21063.2773] [ 4263] [depth= 1] . ( 139 sec) [ -21029.0467] [ 4255] [depth= 1] . ( 139 sec) [ -20978.7661] [ 4238] [depth= 2] . ( 139 sec) [ -20977.8234] [ 4239] [depth= 2] . ( 140 sec) [ -20931.3195] [ 4217] [depth= 5] . ( 140 sec) [ -20719.9400] [ 4122] [depth= 26] . ( 142 sec) [ -20718.9156] [ 4124] [depth= 1] . ( 142 sec) [ -20715.0633] [ 4126] [depth= 1] . ( 143 sec) [ -20712.2920] [ 4124] [depth= 1] . ( 143 sec) [ -20712.2616] [ 4126] [depth= 1] . ( 145 sec) [ -20711.7840] [ 4125] [depth= 1] . ( 146 sec) [ -20711.7668] [ 4123] [depth= 8] . ( 146 sec) [ -20711.3040] [ 4123] [depth= 1] . ( 146 sec) [ -20711.2755] [ 4123] [depth= 1] . ( 147 sec) [ -20711.2420] [ 4123] [depth= 1] . ( 147 sec) [ -20710.9472] [ 4121] [depth= 1] . ( 148 sec) [ -20710.7157] [ 4119] [depth= 1] . ( 148 sec) [ -20703.2547] [ 4118] [depth= 1] . ( 148 sec) [ -20700.5866] [ 4114] [depth= 1] . ( 148 sec) [ -20693.6255] [ 4117] [depth= 1] . ( 149 sec) [ -20691.5347] [ 4115] [depth= 1] . ( 150 sec) [ -20691.2249] [ 4116] [depth= 1] . ( 151 sec) [ -20689.8594] [ 4115] [depth= 2] . ( 152 sec) [ -20689.8258] [ 4120] [depth= 2] . ( 152 sec) [ -20675.0486] [ 4094] [depth= 3] . ( 154 sec) [ -20672.7696] [ 4098] [depth= 1] . ( 155 sec) [ -20671.9245] [ 4099] [depth= 1] . ( 155 sec) [ -20670.6845] [ 4097] [depth= 4] . ( 156 sec) [ -20670.1181] [ 4098] [depth= 1] . ( 157 sec) [ -20667.3381] [ 4101] [depth= 2] . ( 161 sec) [ -20666.3951] [ 4101] [depth= 1] . ( 162 sec) [ -20650.0758] [ 4101] [depth= 2] . ( 163 sec) [ -20636.0556] [ 4092] [depth= 1] . ( 164 sec) [ -20611.1430] [ 4087] [depth= 15] . ( 168 sec) [ -20610.3083] [ 4084] [depth= 1] . ( 169 sec) [ -20607.9195] [ 4083] [depth= 1] . ( 169 sec) [ -20598.2793] [ 4081] [depth= 2] . ( 172 sec) [ -20595.0665] [ 4076] [depth= 4] . ( 176 sec) [ -20590.0303] [ 4074] [depth= 4] . ( 176 sec) [ -20587.2534] [ 4074] [depth= 2] . ( 177 sec) [ -20579.2148] [ 4072] [depth= 1] . ( 177 sec) [ -20579.0379] [ 4070] [depth= 1] . ( 178 sec) [ -20579.0150] [ 4070] [depth= 1] . ( 180 sec) [ -20575.0909] [ 4072] [depth= 1] . ( 181 sec) [ -20569.0923] [ 4069] [depth= 3] . ( 182 sec) [ -20566.9730] [ 4067] [depth= 3] . ( 184 sec) [ -20566.6452] [ 4067] [depth= 1] . ( 184 sec) [ -20565.0630] [ 4067] [depth= 1] . ( 185 sec) [ -20564.5847] [ 4067] [depth= 1] . ( 185 sec) [ -20564.1094] [ 4067] [depth= 1] . ( 186 sec) [ -20564.0461] [ 4069] [depth= 2] . ( 187 sec) [ -20514.5569] [ 4059] [depth= 4] . ( 188 sec) [ -20513.9438] [ 4059] [depth= 1] . ( 188 sec) [ -20512.3204] [ 4058] [depth= 1] . ( 192 sec) [ -20510.9725] [ 4059] [depth= 1] . ( 195 sec) [ -20509.2835] [ 4058] [depth= 6] . ( 195 sec) [ -20506.3473] [ 4057] [depth= 10] . ( 202 sec) [ -20503.5100] [ 4055] [depth= 1] . ( 202 sec) [ -20503.1607] [ 4055] [depth= 1] . ( 203 sec) [ -20499.9933] [ 4055] [depth= 1] . ( 205 sec) [ -20499.8139] [ 4054] [depth= 1] . ( 206 sec) [ -20499.0626] [ 4057] [depth= 1] . ( 206 sec) [ -20498.5006] [ 4058] [depth= 1] . ( 206 sec) [ -20498.3531] [ 4058] [depth= 1] . ( 209 sec) [ -20494.8181] [ 4056] [depth= 2] . ( 209 sec) [ -20493.7499] [ 4056] [depth= 1] . ( 209 sec) [ -20493.5972] [ 4057] [depth= 1] . ( 214 sec) [ -20491.8538] [ 4057] [depth= 1] . ( 214 sec) [ -20491.5955] [ 4057] [depth= 1] . ( 215 sec) [ -20491.4733] [ 4057] [depth= 1] . ( 217 sec) [ -20443.2439] [Branch lengths ] . ( 221 sec) [ -20442.8238] [GTR parameters ] . ( 221 sec) [ -20442.6174] [Alpha ][ 0.340936] . ( 228 sec) [ -20439.9685] [ 4054] [depth= 1] . ( 241 sec) [ -20439.3894] [ 4054] [depth= 1] . ( 244 sec) [ -20430.0119] [ 4052] [depth= 5] . ( 253 sec) [ -20429.7809] [ 4056] [depth= 2] . ( 254 sec) [ -20425.1715] [ 4052] [depth= 3] . ( 255 sec) [ -20425.0794] [ 4052] [depth= 1] . ( 255 sec) [ -20424.9378] [ 4053] [depth= 1] . ( 256 sec) [ -20424.7370] [ 4057] [depth= 5] . ( 259 sec) [ -20424.2167] [ 4058] [depth= 1] . ( 264 sec) [ -20423.8092] [ 4060] [depth= 1] . ( 267 sec) [ -20423.4371] [ 4043] [depth= 2] . ( 269 sec) [ -20422.2014] [ 4040] [depth= 1] . ( 270 sec) [ -20421.7856] [ 4040] [depth= 2] . ( 279 sec) [ -20421.6046] [ 4042] [depth= 4] . ( 285 sec) [ -20420.9370] [ 4042] [depth= 3] . ( 291 sec) [ -20420.8269] [ 4042] [depth= 2] . ( 292 sec) [ -20420.3063] [ 4042] [depth= 5] . ( 301 sec) [ -20419.8033] [ 4041] [depth= 1] . ( 303 sec) [ -20419.5484] [ 4046] [depth= 3] . ( 308 sec) [ -20419.2561] [ 4046] [depth= 1] . ( 326 sec) [ -20419.2236] [ 4046] [depth= 1] . ( 338 sec) [ -20408.9664] [Branch lengths ] . ( 341 sec) [ -20408.8185] [GTR parameters ] . ( 341 sec) [ -20408.8170] [Alpha ][ 0.341507] . ( 351 sec) [ -20408.7207] [ 4049] [depth= 1] . ( 417 sec) [ -20408.6783] [ 4049] [depth= 1] . ( 425 sec) [ -20408.0461] [ 4049] [depth= 3] . ( 437 sec) [ -20407.7154] [ 4053] [depth= 3] . ( 456 sec) [ -20407.6831] [ 4053] [depth= 1] . ( 472 sec) [ -20406.5846] [Branch lengths ] . ( 476 sec) [ -20406.5592] [GTR parameters ] . ( 476 sec) [ -20406.5592] [Alpha ][ 0.341507] . ( 565 sec) [ -20406.4768] [ 4053] [depth= 11] . ( 576 sec) [ -20406.3229] [ 4053] [depth= 1] . ( 594 sec) [ -20406.1417] [ 4052] [depth= 1] . ( 606 sec) [ -20405.3530] [Branch lengths ] . ( 610 sec) [ -20405.3349] [GTR parameters ] . ( 610 sec) [ -20405.3329] [Alpha ][ 0.340655] . ( 737 sec) [ -20405.2859] [Branch lengths ] . ( 851 sec) [ -20405.2716] [ 4059] [depth= 6] . ( 990 sec) [ -20404.6248] [Branch lengths ] . ( 993 sec) [ -20404.6140] [GTR parameters ] . ( 994 sec) [ -20404.6134] [Alpha ][ 0.340122] . ( 994 sec) [ -20404.6134] [Topology ] . ( 999 sec) [ -20404.0578] [Topology ] . ( 1005 sec) [ -20403.8304] [Topology ] . ( 1010 sec) [ -20403.7778] [Topology ] . ( 1015 sec) [ -20403.7600] [Topology ] . ( 1020 sec) [ -20403.7538] [Topology ] . ( 1026 sec) [ -20403.7517] [Topology ] . ( 1031 sec) [ -20403.7510] [Topology ] . ( 1034 sec) [ -20403.7497] [GTR parameters ] . ( 1035 sec) [ -20403.7495] [Alpha ][ 0.340462] . ( 1035 sec) [ -20403.7495] [Topology ] . ( 1040 sec) [ -20403.7492] [Topology ] . ( 1041 sec) [ -20403.7492] [GTR parameters ] . ( 1041 sec) [ -20403.7492] [Alpha ][ 0.340462] . Checking for NNIs, optimizing five branches... . Log likelihood of the current tree: -20403.749201. . [Random start 3/ 5] . Computing pairwise distances... . Building BioNJ tree... . Randomising the tree... . Maximizing likelihood (using SPR moves)... . ( 6 sec) [ -44654.8979] [GTR parameters ] . ( 7 sec) [ -43627.5036] [Alpha ][ 0.500960] . ( 7 sec) [ -43518.5348] [ 7529] [depth= 9] . ( 7 sec) [ -43435.3684] [ 7517] [depth= 14] . ( 7 sec) [ -43411.8042] [ 7512] [depth= 3] . ( 8 sec) [ -43406.9713] [ 7511] [depth= 9] . ( 8 sec) [ -43401.6606] [ 7507] [depth= 2] . ( 8 sec) [ -43293.1603] [ 7507] [depth= 2] . ( 8 sec) [ -43089.9550] [ 7502] [depth= 7] . ( 9 sec) [ -43086.1792] [ 7500] [depth= 6] . ( 9 sec) [ -43059.6454] [ 7497] [depth= 10] . ( 9 sec) [ -43003.0126] [ 7498] [depth= 1] . ( 9 sec) [ -42982.5291] [ 7489] [depth= 5] . ( 9 sec) [ -42946.1359] [ 7487] [depth= 3] . ( 10 sec) [ -42473.2809] [ 7480] [depth= 2] . ( 10 sec) [ -41960.2656] [ 7478] [depth= 25] . ( 10 sec) [ -41867.8064] [ 7467] [depth= 24] . ( 11 sec) [ -41851.6770] [ 7460] [depth= 1] . ( 11 sec) [ -41358.4370] [ 7455] [depth= 6] . ( 11 sec) [ -41208.2847] [ 7447] [depth= 10] . ( 11 sec) [ -41127.3980] [ 7433] [depth= 28] . ( 12 sec) [ -41061.8830] [ 7420] [depth= 23] . ( 12 sec) [ -40983.6751] [ 7404] [depth= 6] . ( 12 sec) [ -40915.0573] [ 7393] [depth= 2] . ( 12 sec) [ -40528.2560] [ 7311] [depth= 20] . ( 13 sec) [ -40523.2570] [ 7302] [depth= 13] . ( 13 sec) [ -39978.5598] [ 7294] [depth= 19] . ( 13 sec) [ -39710.3761] [ 7234] [depth= 17] . ( 13 sec) [ -39700.0395] [ 7194] [depth= 8] . ( 14 sec) [ -39601.2002] [ 7176] [depth= 8] . ( 14 sec) [ -39528.9934] [ 7162] [depth= 4] . ( 14 sec) [ -39487.1216] [ 7149] [depth= 2] . ( 14 sec) [ -39427.3056] [ 7143] [depth= 3] . ( 15 sec) [ -39082.0634] [ 7139] [depth= 25] . ( 15 sec) [ -38439.4145] [ 7137] [depth= 1] . ( 15 sec) [ -37818.4691] [ 7029] [depth= 7] . ( 16 sec) [ -37154.9503] [ 6898] [depth= 5] . ( 16 sec) [ -36982.6383] [ 6871] [depth= 24] . ( 16 sec) [ -36910.5480] [ 6874] [depth= 1] . ( 16 sec) [ -36794.9518] [ 6861] [depth= 9] . ( 17 sec) [ -36177.7601] [ 6850] [depth= 5] . ( 17 sec) [ -35846.1957] [ 6812] [depth= 10] . ( 17 sec) [ -35798.7851] [ 6808] [depth= 8] . ( 18 sec) [ -35721.0947] [ 6795] [depth= 10] . ( 18 sec) [ -35717.0556] [ 6794] [depth= 2] . ( 18 sec) [ -35701.7802] [ 6794] [depth= 1] . ( 18 sec) [ -35671.9713] [ 6780] [depth= 1] . ( 18 sec) [ -35033.5782] [ 6630] [depth= 25] . ( 19 sec) [ -34859.5694] [ 6611] [depth= 10] . ( 19 sec) [ -34541.3482] [ 6581] [depth= 8] . ( 19 sec) [ -34528.9421] [ 6581] [depth= 1] . ( 20 sec) [ -34430.2393] [ 6571] [depth= 1] . ( 20 sec) [ -34106.2428] [ 6562] [depth= 4] . ( 20 sec) [ -34097.8044] [ 6563] [depth= 1] . ( 21 sec) [ -33695.5766] [ 6473] [depth= 2] . ( 21 sec) [ -33528.8933] [ 6412] [depth= 9] . ( 21 sec) [ -33431.1303] [ 6406] [depth= 21] . ( 21 sec) [ -33322.7789] [ 6340] [depth= 5] . ( 22 sec) [ -33130.9726] [ 6285] [depth= 9] . ( 22 sec) [ -32812.9372] [ 6260] [depth= 7] . ( 22 sec) [ -32738.7605] [ 6243] [depth= 2] . ( 23 sec) [ -32710.6597] [ 6240] [depth= 4] . ( 23 sec) [ -32487.7703] [ 6223] [depth= 17] . ( 23 sec) [ -32362.7837] [ 6215] [depth= 7] . ( 23 sec) [ -32358.3299] [ 6212] [depth= 9] . ( 24 sec) [ -32324.5481] [ 6202] [depth= 1] . ( 24 sec) [ -32202.7582] [ 6179] [depth= 10] . ( 24 sec) [ -32123.0405] [ 6163] [depth= 5] . ( 24 sec) [ -32106.5545] [ 6151] [depth= 5] . ( 25 sec) [ -32100.2467] [ 6148] [depth= 11] . ( 25 sec) [ -32060.4674] [ 6143] [depth= 2] . ( 25 sec) [ -32030.5111] [ 6136] [depth= 7] . ( 25 sec) [ -31992.4814] [ 6119] [depth= 13] . ( 26 sec) [ -31669.6304] [ 6116] [depth= 1] . ( 26 sec) [ -31529.1152] [ 6029] [depth= 9] . ( 26 sec) [ -31030.8901] [ 6028] [depth= 3] . ( 26 sec) [ -30591.0160] [ 6025] [depth= 13] . ( 27 sec) [ -30463.2330] [ 5999] [depth= 8] . ( 27 sec) [ -30452.4911] [ 5994] [depth= 4] . ( 27 sec) [ -30435.7612] [ 5992] [depth= 4] . ( 27 sec) [ -30379.8916] [ 5980] [depth= 2] . ( 28 sec) [ -30361.6388] [ 5981] [depth= 1] . ( 28 sec) [ -30314.5226] [ 5972] [depth= 2] . ( 28 sec) [ -30270.4816] [ 5966] [depth= 13] . ( 29 sec) [ -30189.8363] [ 5949] [depth= 25] . ( 29 sec) [ -30116.2066] [ 5939] [depth= 14] . ( 29 sec) [ -29988.2128] [ 5914] [depth= 8] . ( 30 sec) [ -29966.2183] [ 5914] [depth= 10] . ( 30 sec) [ -29928.1496] [ 5922] [depth= 2] . ( 30 sec) [ -29844.2957] [ 5897] [depth= 12] . ( 30 sec) [ -29836.9208] [ 5896] [depth= 1] . ( 31 sec) [ -29819.2533] [ 5886] [depth= 5] . ( 31 sec) [ -29689.7726] [ 5884] [depth= 8] . ( 31 sec) [ -29448.1023] [ 5885] [depth= 19] . ( 31 sec) [ -29383.4752] [ 5911] [depth= 11] . ( 32 sec) [ -29305.9938] [ 5883] [depth= 4] . ( 32 sec) [ -29270.6525] [ 5875] [depth= 18] . ( 32 sec) [ -29236.9565] [ 5871] [depth= 19] . ( 33 sec) [ -29234.4567] [ 5875] [depth= 1] . ( 33 sec) [ -29011.5334] [ 5863] [depth= 12] . ( 33 sec) [ -28949.8063] [ 5847] [depth= 19] . ( 34 sec) [ -28939.1770] [ 5843] [depth= 1] . ( 34 sec) [ -28862.9333] [ 5833] [depth= 3] . ( 34 sec) [ -28859.9637] [ 5833] [depth= 1] . ( 34 sec) [ -28859.6442] [ 5835] [depth= 1] . ( 35 sec) [ -28837.7196] [ 5831] [depth= 14] . ( 35 sec) [ -28818.5545] [ 5826] [depth= 7] . ( 35 sec) [ -28813.3579] [ 5829] [depth= 1] . ( 36 sec) [ -28811.5579] [ 5829] [depth= 1] . ( 36 sec) [ -28762.5318] [ 5815] [depth= 2] . ( 36 sec) [ -28758.7397] [ 5816] [depth= 2] . ( 37 sec) [ -28693.4431] [ 5811] [depth= 5] . ( 37 sec) [ -28641.8858] [ 5799] [depth= 10] . ( 37 sec) [ -28611.9016] [ 5785] [depth= 9] . ( 37 sec) [ -28599.7178] [ 5780] [depth= 1] . ( 38 sec) [ -28595.2739] [ 5782] [depth= 1] . ( 38 sec) [ -28594.1157] [ 5782] [depth= 1] . ( 38 sec) [ -28558.4303] [ 5782] [depth= 14] . ( 38 sec) [ -28414.5809] [ 5772] [depth= 11] . ( 39 sec) [ -28408.7442] [ 5770] [depth= 1] . ( 39 sec) [ -28394.7610] [ 5762] [depth= 9] . ( 39 sec) [ -28380.3428] [ 5758] [depth= 4] . ( 40 sec) [ -28378.7560] [ 5760] [depth= 1] . ( 40 sec) [ -28346.3667] [ 5759] [depth= 2] . ( 40 sec) [ -28330.0978] [ 5751] [depth= 8] . ( 40 sec) [ -28329.3734] [ 5745] [depth= 8] . ( 41 sec) [ -28290.8305] [ 5740] [depth= 7] . ( 41 sec) [ -28282.4618] [ 5737] [depth= 2] . ( 41 sec) [ -28231.3414] [ 5724] [depth= 8] . ( 41 sec) [ -28231.1269] [ 5726] [depth= 1] . ( 42 sec) [ -28210.9819] [ 5730] [depth= 1] . ( 42 sec) [ -28063.5608] [ 5725] [depth= 8] . ( 42 sec) [ -27993.6909] [ 5699] [depth= 16] . ( 43 sec) [ -27974.9088] [ 5695] [depth= 7] . ( 43 sec) [ -27970.7368] [ 5695] [depth= 1] . ( 43 sec) [ -27968.2356] [ 5693] [depth= 2] . ( 43 sec) [ -27950.0444] [ 5695] [depth= 6] . ( 44 sec) [ -27941.2543] [ 5691] [depth= 11] . ( 44 sec) [ -27881.8174] [ 5676] [depth= 6] . ( 44 sec) [ -27879.4148] [ 5679] [depth= 5] . ( 45 sec) [ -27862.0259] [ 5677] [depth= 2] . ( 45 sec) [ -27860.6507] [ 5677] [depth= 1] . ( 45 sec) [ -27832.4854] [ 5678] [depth= 1] . ( 45 sec) [ -27769.9541] [ 5675] [depth= 14] . ( 46 sec) [ -27731.1842] [ 5674] [depth= 4] . ( 46 sec) [ -27468.1846] [ 5655] [depth= 1] . ( 46 sec) [ -27426.1588] [ 5652] [depth= 20] . ( 46 sec) [ -27425.8537] [ 5651] [depth= 2] . ( 47 sec) [ -27370.8553] [ 5641] [depth= 3] . ( 47 sec) [ -27346.5521] [ 5635] [depth= 9] . ( 47 sec) [ -27344.6931] [ 5631] [depth= 5] . ( 48 sec) [ -27325.4973] [ 5625] [depth= 14] . ( 48 sec) [ -27303.2156] [ 5620] [depth= 10] . ( 48 sec) [ -27296.3232] [ 5625] [depth= 20] . ( 49 sec) [ -27289.7765] [ 5622] [depth= 13] . ( 49 sec) [ -27137.7797] [ 5532] [depth= 1] . ( 49 sec) [ -27120.4451] [ 5531] [depth= 15] . ( 50 sec) [ -27086.9192] [ 5524] [depth= 18] . ( 50 sec) [ -26850.0236] [ 5484] [depth= 22] . ( 50 sec) [ -26766.4811] [ 5461] [depth= 19] . ( 50 sec) [ -26752.9721] [ 5460] [depth= 2] . ( 51 sec) [ -26735.1370] [ 5456] [depth= 5] . ( 51 sec) [ -26690.5576] [ 5449] [depth= 21] . ( 51 sec) [ -26624.1292] [ 5449] [depth= 1] . ( 52 sec) [ -26559.5964] [ 5422] [depth= 26] . ( 52 sec) [ -26536.6757] [ 5418] [depth= 1] . ( 52 sec) [ -26469.1517] [ 5407] [depth= 1] . ( 52 sec) [ -26459.2870] [ 5404] [depth= 13] . ( 53 sec) [ -26430.4278] [ 5401] [depth= 1] . ( 53 sec) [ -26401.7266] [ 5401] [depth= 8] . ( 53 sec) [ -26383.8594] [ 5404] [depth= 15] . ( 54 sec) [ -26297.9887] [ 5382] [depth= 15] . ( 54 sec) [ -26277.6656] [ 5377] [depth= 1] . ( 54 sec) [ -26116.8969] [ 5343] [depth= 22] . ( 55 sec) [ -25999.5310] [ 5334] [depth= 11] . ( 55 sec) [ -25973.9862] [ 5330] [depth= 4] . ( 55 sec) [ -25947.4911] [ 5320] [depth= 4] . ( 55 sec) [ -25892.5803] [ 5316] [depth= 5] . ( 55 sec) [ -25859.9921] [ 5290] [depth= 1] . ( 56 sec) [ -25838.0715] [ 5290] [depth= 21] . ( 56 sec) [ -25678.1403] [ 5269] [depth= 8] . ( 56 sec) [ -25500.2625] [ 5265] [depth= 3] . ( 57 sec) [ -25379.8158] [ 5206] [depth= 9] . ( 57 sec) [ -25352.0128] [ 5196] [depth= 1] . ( 57 sec) [ -25306.1926] [ 5186] [depth= 19] . ( 58 sec) [ -25301.9816] [ 5184] [depth= 1] . ( 58 sec) [ -25284.5173] [ 5180] [depth= 16] . ( 58 sec) [ -25195.3342] [ 5160] [depth= 19] . ( 59 sec) [ -24753.5878] [ 5059] [depth= 13] . ( 60 sec) [ -24326.7427] [ 5014] [depth= 24] . ( 60 sec) [ -24260.8869] [ 4971] [depth= 8] . ( 60 sec) [ -24230.9148] [ 4958] [depth= 22] . ( 61 sec) [ -24217.5739] [ 4954] [depth= 18] . ( 61 sec) [ -24050.4401] [ 4921] [depth= 18] . ( 61 sec) [ -23984.1918] [ 4910] [depth= 16] . ( 61 sec) [ -23946.7008] [ 4912] [depth= 1] . ( 62 sec) [ -23937.7849] [ 4910] [depth= 1] . ( 62 sec) [ -23834.7426] [ 4882] [depth= 16] . ( 62 sec) [ -23822.2373] [ 4880] [depth= 1] . ( 63 sec) [ -23667.9623] [ 4863] [depth= 15] . ( 63 sec) [ -23642.6429] [ 4857] [depth= 2] . ( 63 sec) [ -23604.3757] [ 4844] [depth= 3] . ( 64 sec) [ -23580.8121] [ 4842] [depth= 21] . ( 64 sec) [ -23561.7460] [ 4832] [depth= 3] . ( 64 sec) [ -23412.5459] [ 4813] [depth= 3] . ( 65 sec) [ -23410.9600] [ 4810] [depth= 3] . ( 65 sec) [ -23399.1129] [ 4803] [depth= 1] . ( 65 sec) [ -23379.0780] [ 4803] [depth= 13] . ( 66 sec) [ -23367.0981] [ 4800] [depth= 10] . ( 66 sec) [ -23364.1559] [ 4801] [depth= 2] . ( 66 sec) [ -23364.1430] [ 4802] [depth= 5] . ( 66 sec) [ -23351.4733] [ 4802] [depth= 1] . ( 66 sec) [ -23349.4542] [ 4802] [depth= 1] . ( 67 sec) [ -23320.4445] [ 4796] [depth= 5] . ( 67 sec) [ -23314.7683] [ 4793] [depth= 12] . ( 67 sec) [ -23241.0739] [ 4764] [depth= 2] . ( 67 sec) [ -23229.1742] [ 4749] [depth= 13] . ( 68 sec) [ -23185.7089] [ 4739] [depth= 15] . ( 68 sec) [ -23183.6811] [ 4741] [depth= 1] . ( 68 sec) [ -23183.0142] [ 4743] [depth= 1] . ( 69 sec) [ -23160.9686] [ 4740] [depth= 7] . ( 69 sec) [ -23151.3179] [ 4736] [depth= 13] . ( 69 sec) [ -23148.0900] [ 4735] [depth= 2] . ( 70 sec) [ -23146.3192] [ 4736] [depth= 1] . ( 70 sec) [ -23096.8451] [ 4726] [depth= 11] . ( 70 sec) [ -23095.8494] [ 4727] [depth= 1] . ( 70 sec) [ -23093.7127] [ 4728] [depth= 16] . ( 71 sec) [ -23085.4978] [ 4727] [depth= 1] . ( 71 sec) [ -23085.1913] [ 4727] [depth= 1] . ( 71 sec) [ -23029.5538] [ 4710] [depth= 13] . ( 72 sec) [ -23027.1890] [ 4709] [depth= 1] . ( 72 sec) [ -23017.9065] [ 4707] [depth= 14] . ( 72 sec) [ -23010.3114] [ 4705] [depth= 5] . ( 73 sec) [ -23008.4125] [ 4705] [depth= 1] . ( 73 sec) [ -22942.8243] [ 4689] [depth= 4] . ( 74 sec) [ -22935.9921] [ 4686] [depth= 14] . ( 75 sec) [ -22933.6304] [ 4686] [depth= 1] . ( 75 sec) [ -22925.1070] [ 4681] [depth= 4] . ( 75 sec) [ -22874.8056] [ 4682] [depth= 1] . ( 76 sec) [ -22871.4153] [ 4681] [depth= 3] . ( 76 sec) [ -22842.2440] [ 4674] [depth= 4] . ( 76 sec) [ -22787.8734] [ 4664] [depth= 9] . ( 76 sec) [ -22783.5604] [ 4664] [depth= 2] . ( 79 sec) [ -22782.4856] [ 4664] [depth= 1] . ( 79 sec) [ -22744.9015] [ 4653] [depth= 2] . ( 80 sec) [ -22723.6198] [ 4646] [depth= 11] . ( 80 sec) [ -22720.6478] [ 4645] [depth= 1] . ( 80 sec) [ -22713.9109] [ 4643] [depth= 1] . ( 80 sec) [ -22608.2962] [ 4617] [depth= 3] . ( 81 sec) [ -22596.0980] [ 4612] [depth= 7] . ( 81 sec) [ -22593.5899] [ 4612] [depth= 1] . ( 81 sec) [ -22538.0255] [ 4593] [depth= 3] . ( 82 sec) [ -22512.7547] [ 4592] [depth= 1] . ( 82 sec) [ -22512.0659] [ 4593] [depth= 2] . ( 82 sec) [ -22508.1032] [ 4591] [depth= 1] . ( 82 sec) [ -22491.4228] [ 4589] [depth= 7] . ( 83 sec) [ -22484.8971] [ 4588] [depth= 9] . ( 83 sec) [ -22462.2748] [ 4588] [depth= 10] . ( 83 sec) [ -22321.5895] [ 4550] [depth= 1] . ( 83 sec) [ -22320.2734] [ 4551] [depth= 1] . ( 84 sec) [ -22307.3900] [ 4550] [depth= 6] . ( 84 sec) [ -22294.1101] [ 4549] [depth= 1] . ( 84 sec) [ -22230.0613] [ 4503] [depth= 21] . ( 84 sec) [ -22228.7533] [ 4504] [depth= 6] . ( 85 sec) [ -22219.1456] [ 4502] [depth= 9] . ( 85 sec) [ -22202.2403] [ 4499] [depth= 2] . ( 85 sec) [ -22200.3196] [ 4502] [depth= 1] . ( 86 sec) [ -22175.2650] [ 4506] [depth= 18] . ( 87 sec) [ -22168.5331] [ 4495] [depth= 1] . ( 87 sec) [ -22164.7920] [ 4491] [depth= 5] . ( 87 sec) [ -22163.6257] [ 4490] [depth= 1] . ( 88 sec) [ -22157.7776] [ 4489] [depth= 1] . ( 88 sec) [ -22156.6168] [ 4489] [depth= 1] . ( 88 sec) [ -22153.1703] [ 4488] [depth= 3] . ( 89 sec) [ -22144.4539] [ 4486] [depth= 2] . ( 89 sec) [ -22143.3748] [ 4487] [depth= 1] . ( 90 sec) [ -22141.1547] [ 4487] [depth= 3] . ( 90 sec) [ -22124.7322] [ 4478] [depth= 2] . ( 90 sec) [ -22118.8286] [ 4478] [depth= 1] . ( 90 sec) [ -22116.8877] [ 4480] [depth= 3] . ( 91 sec) [ -22105.3311] [ 4474] [depth= 5] . ( 91 sec) [ -22099.9794] [ 4469] [depth= 5] . ( 92 sec) [ -22095.9765] [ 4466] [depth= 12] . ( 92 sec) [ -22091.7147] [ 4464] [depth= 2] . ( 92 sec) [ -22090.4776] [ 4464] [depth= 1] . ( 93 sec) [ -22082.7037] [ 4463] [depth= 1] . ( 93 sec) [ -22079.6419] [ 4461] [depth= 1] . ( 93 sec) [ -22068.3661] [ 4462] [depth= 1] . ( 94 sec) [ -22067.8030] [ 4462] [depth= 1] . ( 94 sec) [ -22064.7643] [ 4462] [depth= 1] . ( 94 sec) [ -22062.9963] [ 4464] [depth= 1] . ( 94 sec) [ -22061.3436] [ 4464] [depth= 1] . ( 95 sec) [ -22027.1102] [ 4447] [depth= 22] . ( 95 sec) [ -21968.8753] [ 4428] [depth= 1] . ( 95 sec) [ -21961.0090] [ 4427] [depth= 1] . ( 96 sec) [ -21955.0627] [ 4425] [depth= 15] . ( 96 sec) [ -21924.1437] [ 4418] [depth= 2] . ( 97 sec) [ -21922.3323] [ 4416] [depth= 2] . ( 97 sec) [ -21918.8526] [ 4415] [depth= 1] . ( 97 sec) [ -21912.9053] [ 4415] [depth= 3] . ( 98 sec) [ -21910.7029] [ 4414] [depth= 10] . ( 98 sec) [ -21909.6466] [ 4412] [depth= 1] . ( 98 sec) [ -21890.9678] [ 4408] [depth= 4] . ( 99 sec) [ -21878.9158] [ 4404] [depth= 10] . ( 99 sec) [ -21872.9978] [ 4403] [depth= 10] . ( 100 sec) [ -21844.5487] [ 4397] [depth= 8] . ( 100 sec) [ -21744.8345] [ 4377] [depth= 1] . ( 101 sec) [ -21736.8250] [ 4371] [depth= 1] . ( 101 sec) [ -21728.8977] [ 4370] [depth= 21] . ( 101 sec) [ -21723.7946] [ 4362] [depth= 14] . ( 102 sec) [ -21714.4299] [ 4360] [depth= 1] . ( 102 sec) [ -21712.8108] [ 4361] [depth= 1] . ( 104 sec) [ -21352.0153] [Branch lengths ] . ( 111 sec) [ -21325.8201] [GTR parameters ] . ( 112 sec) [ -21268.3860] [Alpha ][ 0.342663] . ( 112 sec) [ -21268.3722] [ 4357] [depth= 2] . ( 112 sec) [ -21264.7515] [ 4357] [depth= 1] . ( 113 sec) [ -21263.1139] [ 4351] [depth= 8] . ( 113 sec) [ -21256.2721] [ 4340] [depth= 3] . ( 113 sec) [ -21241.4181] [ 4320] [depth= 20] . ( 114 sec) [ -21236.7070] [ 4316] [depth= 19] . ( 114 sec) [ -21231.8665] [ 4313] [depth= 4] . ( 115 sec) [ -21183.2805] [ 4298] [depth= 13] . ( 115 sec) [ -21180.9101] [ 4296] [depth= 1] . ( 115 sec) [ -21179.5565] [ 4295] [depth= 2] . ( 116 sec) [ -21176.8564] [ 4294] [depth= 1] . ( 116 sec) [ -21165.4775] [ 4294] [depth= 15] . ( 117 sec) [ -21159.2146] [ 4293] [depth= 4] . ( 117 sec) [ -21158.7612] [ 4292] [depth= 1] . ( 118 sec) [ -21154.0435] [ 4291] [depth= 2] . ( 118 sec) [ -21151.1508] [ 4287] [depth= 6] . ( 118 sec) [ -21149.6027] [ 4287] [depth= 1] . ( 119 sec) [ -21148.6597] [ 4285] [depth= 1] . ( 119 sec) [ -21148.6153] [ 4288] [depth= 2] . ( 119 sec) [ -21135.1177] [ 4272] [depth= 11] . ( 120 sec) [ -21126.6267] [ 4276] [depth= 15] . ( 121 sec) [ -21123.8675] [ 4278] [depth= 1] . ( 122 sec) [ -21123.0298] [ 4277] [depth= 1] . ( 123 sec) [ -21119.5881] [ 4275] [depth= 1] . ( 124 sec) [ -21116.1133] [ 4267] [depth= 2] . ( 124 sec) [ -21010.9169] [ 4217] [depth= 7] . ( 125 sec) [ -21009.7988] [ 4218] [depth= 1] . ( 125 sec) [ -21002.9159] [ 4208] [depth= 2] . ( 125 sec) [ -20965.0374] [ 4190] [depth= 14] . ( 126 sec) [ -20947.6321] [ 4186] [depth= 8] . ( 127 sec) [ -20947.3440] [ 4186] [depth= 1] . ( 127 sec) [ -20939.7187] [ 4184] [depth= 1] . ( 127 sec) [ -20939.5629] [ 4183] [depth= 1] . ( 128 sec) [ -20938.4164] [ 4181] [depth= 1] . ( 128 sec) [ -20927.7833] [ 4178] [depth= 1] . ( 128 sec) [ -20915.8071] [ 4176] [depth= 1] . ( 131 sec) [ -20886.9390] [ 4170] [depth= 2] . ( 132 sec) [ -20882.0470] [ 4146] [depth= 15] . ( 134 sec) [ -20881.4759] [ 4144] [depth= 2] . ( 136 sec) [ -20880.9191] [ 4147] [depth= 3] . ( 136 sec) [ -20880.6777] [ 4145] [depth= 1] . ( 136 sec) [ -20879.1953] [ 4145] [depth= 4] . ( 137 sec) [ -20876.3794] [ 4146] [depth= 1] . ( 137 sec) [ -20874.0210] [ 4146] [depth= 1] . ( 141 sec) [ -20871.0925] [ 4147] [depth= 2] . ( 142 sec) [ -20864.6868] [ 4144] [depth= 1] . ( 143 sec) [ -20854.8949] [ 4142] [depth= 3] . ( 146 sec) [ -20852.0678] [ 4143] [depth= 1] . ( 150 sec) [ -20849.2081] [ 4142] [depth= 2] . ( 151 sec) [ -20845.7474] [ 4141] [depth= 1] . ( 152 sec) [ -20844.8615] [ 4140] [depth= 2] . ( 153 sec) [ -20839.9718] [ 4141] [depth= 1] . ( 154 sec) [ -20839.6385] [ 4141] [depth= 1] . ( 155 sec) [ -20831.0295] [ 4137] [depth= 3] . ( 155 sec) [ -20829.8724] [ 4137] [depth= 1] . ( 157 sec) [ -20740.1735] [ 4115] [depth= 13] . ( 158 sec) [ -20730.0159] [ 4115] [depth= 1] . ( 159 sec) [ -20727.6514] [ 4113] [depth= 26] . ( 160 sec) [ -20727.3974] [ 4113] [depth= 1] . ( 160 sec) [ -20724.6160] [ 4114] [depth= 1] . ( 161 sec) [ -20724.1850] [ 4114] [depth= 1] . ( 162 sec) [ -20709.6747] [ 4112] [depth= 11] . ( 163 sec) [ -20709.0138] [ 4113] [depth= 1] . ( 163 sec) [ -20706.2069] [ 4116] [depth= 1] . ( 164 sec) [ -20696.3348] [ 4114] [depth= 9] . ( 164 sec) [ -20694.8749] [ 4115] [depth= 1] . ( 164 sec) [ -20694.7958] [ 4113] [depth= 1] . ( 165 sec) [ -20640.0996] [ 4094] [depth= 7] . ( 166 sec) [ -20637.5727] [ 4093] [depth= 2] . ( 166 sec) [ -20618.9067] [ 4083] [depth= 13] . ( 167 sec) [ -20616.3070] [ 4085] [depth= 2] . ( 167 sec) [ -20615.3109] [ 4086] [depth= 1] . ( 168 sec) [ -20614.8869] [ 4085] [depth= 2] . ( 169 sec) [ -20604.0068] [ 4083] [depth= 4] . ( 169 sec) [ -20592.7104] [ 4084] [depth= 13] . ( 171 sec) [ -20592.6787] [ 4085] [depth= 1] . ( 172 sec) [ -20591.4828] [ 4085] [depth= 1] . ( 172 sec) [ -20589.0851] [ 4083] [depth= 1] . ( 172 sec) [ -20588.8805] [ 4084] [depth= 1] . ( 174 sec) [ -20588.8110] [ 4082] [depth= 1] . ( 174 sec) [ -20586.5797] [ 4082] [depth= 4] . ( 175 sec) [ -20586.4991] [ 4082] [depth= 1] . ( 177 sec) [ -20586.4873] [ 4082] [depth= 1] . ( 178 sec) [ -20585.5996] [ 4079] [depth= 12] . ( 178 sec) [ -20585.5022] [ 4078] [depth= 1] . ( 179 sec) [ -20584.3888] [ 4079] [depth= 3] . ( 181 sec) [ -20584.0713] [ 4078] [depth= 6] . ( 182 sec) [ -20584.0046] [ 4080] [depth= 1] . ( 183 sec) [ -20579.9342] [ 4077] [depth= 4] . ( 183 sec) [ -20578.0761] [ 4076] [depth= 1] . ( 184 sec) [ -20574.7445] [ 4074] [depth= 2] . ( 185 sec) [ -20574.7261] [ 4075] [depth= 1] . ( 186 sec) [ -20573.5344] [ 4075] [depth= 1] . ( 188 sec) [ -20573.0308] [ 4076] [depth= 1] . ( 189 sec) [ -20572.5639] [ 4077] [depth= 1] . ( 190 sec) [ -20571.2540] [ 4076] [depth= 1] . ( 190 sec) [ -20570.0890] [ 4075] [depth= 2] . ( 196 sec) [ -20569.2449] [ 4075] [depth= 1] . ( 197 sec) [ -20568.6505] [ 4074] [depth= 1] . ( 197 sec) [ -20567.6159] [ 4073] [depth= 1] . ( 198 sec) [ -20565.4451] [ 4074] [depth= 1] . ( 199 sec) [ -20561.5097] [ 4074] [depth= 16] . ( 199 sec) [ -20561.3404] [ 4072] [depth= 1] . ( 201 sec) [ -20560.4665] [ 4072] [depth= 2] . ( 202 sec) [ -20555.6617] [ 4070] [depth= 15] . ( 202 sec) [ -20554.3249] [ 4071] [depth= 1] . ( 205 sec) [ -20550.2196] [ 4068] [depth= 3] . ( 210 sec) [ -20549.5703] [ 4070] [depth= 1] . ( 213 sec) [ -20484.7656] [Branch lengths ] . ( 217 sec) [ -20483.9987] [GTR parameters ] . ( 218 sec) [ -20483.9827] [Alpha ][ 0.340438] . ( 220 sec) [ -20483.4984] [ 4069] [depth= 1] . ( 225 sec) [ -20481.6596] [ 4069] [depth= 1] . ( 231 sec) [ -20479.0975] [ 4068] [depth= 6] . ( 231 sec) [ -20476.8767] [ 4070] [depth= 1] . ( 232 sec) [ -20460.8078] [ 4063] [depth= 17] . ( 233 sec) [ -20459.2591] [ 4062] [depth= 1] . ( 233 sec) [ -20456.0345] [ 4061] [depth= 1] . ( 235 sec) [ -20455.6023] [ 4063] [depth= 2] . ( 236 sec) [ -20455.1272] [ 4059] [depth= 2] . ( 236 sec) [ -20455.0315] [ 4059] [depth= 1] . ( 244 sec) [ -20453.6829] [ 4056] [depth= 4] . ( 245 sec) [ -20451.4866] [ 4056] [depth= 1] . ( 247 sec) [ -20451.2681] [ 4055] [depth= 1] . ( 254 sec) [ -20439.9722] [ 4044] [depth= 10] . ( 256 sec) [ -20438.6096] [ 4043] [depth= 2] . ( 258 sec) [ -20438.5868] [ 4043] [depth= 1] . ( 262 sec) [ -20438.4960] [ 4044] [depth= 1] . ( 267 sec) [ -20438.3348] [ 4044] [depth= 1] . ( 269 sec) [ -20438.2899] [ 4044] [depth= 1] . ( 271 sec) [ -20436.9043] [ 4040] [depth= 1] . ( 272 sec) [ -20436.7004] [ 4040] [depth= 1] . ( 273 sec) [ -20436.5870] [ 4040] [depth= 1] . ( 273 sec) [ -20436.3179] [ 4043] [depth= 2] . ( 274 sec) [ -20435.4245] [ 4045] [depth= 1] . ( 274 sec) [ -20434.5992] [ 4043] [depth= 1] . ( 276 sec) [ -20434.5507] [ 4045] [depth= 1] . ( 278 sec) [ -20433.8982] [ 4050] [depth= 2] . ( 280 sec) [ -20433.7508] [ 4051] [depth= 1] . ( 282 sec) [ -20432.5715] [ 4050] [depth= 1] . ( 286 sec) [ -20432.4160] [ 4049] [depth= 1] . ( 296 sec) [ -20432.3459] [ 4048] [depth= 1] . ( 297 sec) [ -20432.0113] [ 4049] [depth= 1] . ( 302 sec) [ -20431.8769] [ 4050] [depth= 1] . ( 307 sec) [ -20431.4902] [ 4050] [depth= 1] . ( 318 sec) [ -20430.6887] [ 4050] [depth= 1] . ( 323 sec) [ -20426.9490] [ 4046] [depth= 1] . ( 325 sec) [ -20426.4787] [ 4046] [depth= 1] . ( 334 sec) [ -20413.9304] [Branch lengths ] . ( 338 sec) [ -20413.8544] [GTR parameters ] . ( 339 sec) [ -20413.8524] [Alpha ][ 0.341117] . ( 365 sec) [ -20410.9823] [ 4043] [depth= 5] . ( 365 sec) [ -20409.9928] [ 4043] [depth= 1] . ( 369 sec) [ -20409.7693] [ 4047] [depth= 1] . ( 388 sec) [ -20409.6127] [ 4048] [depth= 1] . ( 437 sec) [ -20409.5935] [ 4047] [depth= 1] . ( 439 sec) [ -20409.5184] [ 4046] [depth= 1] . ( 444 sec) [ -20407.8098] [ 4047] [depth= 1] . ( 450 sec) [ -20407.7171] [ 4047] [depth= 2] . ( 467 sec) [ -20404.9400] [Branch lengths ] . ( 471 sec) [ -20404.9183] [GTR parameters ] . ( 471 sec) [ -20404.9183] [Alpha ][ 0.341117] . ( 492 sec) [ -20403.4502] [ 4050] [depth= 2] . ( 502 sec) [ -20403.1006] [ 4046] [depth= 3] . ( 506 sec) [ -20403.0255] [ 4045] [depth= 1] . ( 510 sec) [ -20402.9350] [ 4047] [depth= 1] . ( 512 sec) [ -20402.5363] [ 4046] [depth= 1] . ( 601 sec) [ -20400.0628] [Branch lengths ] . ( 605 sec) [ -20400.0431] [GTR parameters ] . ( 605 sec) [ -20400.0414] [Alpha ][ 0.340333] . ( 636 sec) [ -20399.8852] [ 4050] [depth= 3] . ( 735 sec) [ -20399.5709] [Branch lengths ] . ( 738 sec) [ -20399.5669] [GTR parameters ] . ( 739 sec) [ -20399.5667] [Alpha ][ 0.339993] . ( 778 sec) [ -20399.5409] [ 4056] [depth= 1] . ( 989 sec) [ -20399.4859] [Branch lengths ] . ( 990 sec) [ -20399.4859] [GTR parameters ] . ( 991 sec) [ -20399.4859] [Alpha ][ 0.339993] . ( 991 sec) [ -20399.4859] [Topology ] . ( 996 sec) [ -20399.4700] [Topology ] . ( 1001 sec) [ -20399.4617] [Topology ] . ( 1006 sec) [ -20399.4563] [Topology ] . ( 1012 sec) [ -20399.4526] [Topology ] . ( 1017 sec) [ -20399.4501] [Topology ] . ( 1022 sec) [ -20399.4484] [Topology ] . ( 1023 sec) [ -20399.4484] [GTR parameters ] . ( 1023 sec) [ -20399.4484] [Alpha ][ 0.339993] . ( 1024 sec) [ -20399.4484] [Topology ] . ( 1029 sec) [ -20399.4471] [Topology ] . ( 1034 sec) [ -20399.4463] [Topology ] . ( 1035 sec) [ -20399.4462] [GTR parameters ] . ( 1035 sec) [ -20399.4462] [Alpha ][ 0.339993] . ( 1035 sec) [ -20399.4462] [Topology ] . ( 1041 sec) [ -20399.4456] [Topology ] . ( 1042 sec) [ -20399.4456] [GTR parameters ] . ( 1042 sec) [ -20399.4456] [Alpha ][ 0.339993] . Checking for NNIs, optimizing five branches... . ( 1080 sec) [ -20399.4132] [Topology ] . Checking for NNIs, optimizing five branches... . ( 1117 sec) [ -20399.3966] [Topology ] . Checking for NNIs, optimizing five branches... . ( 1155 sec) [ -20399.3965] [Topology ] . Checking for NNIs, optimizing five branches... . Log likelihood of the current tree: -20399.396470. . [Random start 4/ 5] . Computing pairwise distances... . Building BioNJ tree... . Randomising the tree... . Maximizing likelihood (using SPR moves)... . ( 6 sec) [ -42550.2217] [GTR parameters ] . ( 7 sec) [ -41545.3734] [Alpha ][ 0.500960] . ( 7 sec) [ -41401.3134] [ 7562] [depth= 14] . ( 7 sec) [ -41301.4104] [ 7545] [depth= 13] . ( 8 sec) [ -41035.9591] [ 7531] [depth= 2] . ( 8 sec) [ -41026.0541] [ 7529] [depth= 6] . ( 8 sec) [ -40955.8843] [ 7520] [depth= 1] . ( 8 sec) [ -40909.4075] [ 7509] [depth= 15] . ( 8 sec) [ -40721.1032] [ 7505] [depth= 3] . ( 9 sec) [ -40715.2834] [ 7497] [depth= 12] . ( 9 sec) [ -40695.1315] [ 7478] [depth= 3] . ( 9 sec) [ -40673.1479] [ 7477] [depth= 11] . ( 9 sec) [ -40656.0314] [ 7473] [depth= 6] . ( 10 sec) [ -40649.0899] [ 7470] [depth= 4] . ( 10 sec) [ -40521.3175] [ 7463] [depth= 6] . ( 10 sec) [ -40460.2212] [ 7453] [depth= 11] . ( 10 sec) [ -40449.0135] [ 7452] [depth= 4] . ( 10 sec) [ -40441.4679] [ 7452] [depth= 5] . ( 11 sec) [ -40433.9564] [ 7452] [depth= 6] . ( 11 sec) [ -40390.4916] [ 7452] [depth= 1] . ( 11 sec) [ -40325.3721] [ 7445] [depth= 5] . ( 11 sec) [ -40269.4946] [ 7432] [depth= 9] . ( 12 sec) [ -40197.8520] [ 7421] [depth= 5] . ( 12 sec) [ -40171.5181] [ 7416] [depth= 11] . ( 12 sec) [ -40116.7430] [ 7414] [depth= 19] . ( 12 sec) [ -40095.7847] [ 7412] [depth= 9] . ( 13 sec) [ -39988.5493] [ 7397] [depth= 20] . ( 13 sec) [ -39962.3094] [ 7394] [depth= 5] . ( 13 sec) [ -39940.0688] [ 7393] [depth= 6] . ( 13 sec) [ -39809.8413] [ 7383] [depth= 25] . ( 14 sec) [ -39785.2496] [ 7382] [depth= 1] . ( 14 sec) [ -39588.4185] [ 7366] [depth= 25] . ( 14 sec) [ -39352.9658] [ 7333] [depth= 25] . ( 14 sec) [ -39267.0937] [ 7300] [depth= 1] . ( 15 sec) [ -39066.8400] [ 7257] [depth= 24] . ( 15 sec) [ -39029.8793] [ 7250] [depth= 21] . ( 15 sec) [ -39025.9693] [ 7249] [depth= 3] . ( 16 sec) [ -38896.1641] [ 7215] [depth= 13] . ( 16 sec) [ -38833.3981] [ 7212] [depth= 7] . ( 16 sec) [ -38779.5541] [ 7209] [depth= 4] . ( 16 sec) [ -38777.8374] [ 7213] [depth= 2] . ( 17 sec) [ -38622.5776] [ 7193] [depth= 3] . ( 17 sec) [ -38576.8744] [ 7188] [depth= 13] . ( 17 sec) [ -38572.8225] [ 7183] [depth= 23] . ( 17 sec) [ -38509.5984] [ 7182] [depth= 9] . ( 18 sec) [ -38491.0235] [ 7181] [depth= 7] . ( 18 sec) [ -38251.5848] [ 7147] [depth= 7] . ( 18 sec) [ -37937.6535] [ 7091] [depth= 25] . ( 18 sec) [ -37651.6179] [ 7107] [depth= 8] . ( 19 sec) [ -37573.6398] [ 7099] [depth= 20] . ( 19 sec) [ -37565.4992] [ 7097] [depth= 1] . ( 19 sec) [ -37245.0027] [ 7024] [depth= 16] . ( 19 sec) [ -37194.7586] [ 7021] [depth= 8] . ( 20 sec) [ -37191.1130] [ 7021] [depth= 1] . ( 20 sec) [ -37047.6937] [ 7003] [depth= 27] . ( 20 sec) [ -36941.4963] [ 7004] [depth= 2] . ( 20 sec) [ -36915.8062] [ 7002] [depth= 1] . ( 21 sec) [ -36723.1083] [ 6996] [depth= 21] . ( 21 sec) [ -36712.0457] [ 6994] [depth= 1] . ( 21 sec) [ -36651.9622] [ 6981] [depth= 21] . ( 21 sec) [ -36446.3557] [ 6948] [depth= 8] . ( 22 sec) [ -36025.6177] [ 6934] [depth= 24] . ( 22 sec) [ -35851.5744] [ 6909] [depth= 18] . ( 22 sec) [ -35500.7946] [ 6907] [depth= 25] . ( 23 sec) [ -35248.7987] [ 6894] [depth= 2] . ( 23 sec) [ -35222.6166] [ 6884] [depth= 18] . ( 23 sec) [ -34461.2464] [ 6725] [depth= 12] . ( 23 sec) [ -33992.0268] [ 6570] [depth= 15] . ( 24 sec) [ -33966.5840] [ 6573] [depth= 1] . ( 24 sec) [ -33880.8030] [ 6564] [depth= 14] . ( 24 sec) [ -33879.0249] [ 6560] [depth= 3] . ( 24 sec) [ -33845.0059] [ 6554] [depth= 23] . ( 24 sec) [ -33720.1865] [ 6545] [depth= 2] . ( 25 sec) [ -33681.1695] [ 6535] [depth= 7] . ( 25 sec) [ -33642.5530] [ 6533] [depth= 8] . ( 25 sec) [ -33637.9555] [ 6531] [depth= 4] . ( 25 sec) [ -33628.4994] [ 6525] [depth= 9] . ( 26 sec) [ -33090.2370] [ 6520] [depth= 7] . ( 26 sec) [ -33007.9874] [ 6520] [depth= 2] . ( 26 sec) [ -32985.0815] [ 6524] [depth= 6] . ( 26 sec) [ -32809.9931] [ 6504] [depth= 30] . ( 27 sec) [ -32762.9237] [ 6491] [depth= 8] . ( 27 sec) [ -32739.6989] [ 6484] [depth= 2] . ( 27 sec) [ -32701.5478] [ 6483] [depth= 1] . ( 27 sec) [ -32240.6087] [ 6472] [depth= 19] . ( 28 sec) [ -32207.4800] [ 6468] [depth= 9] . ( 28 sec) [ -32197.5492] [ 6464] [depth= 6] . ( 28 sec) [ -32079.1179] [ 6458] [depth= 13] . ( 28 sec) [ -32074.4168] [ 6457] [depth= 3] . ( 29 sec) [ -31729.5159] [ 6376] [depth= 12] . ( 29 sec) [ -31718.6959] [ 6372] [depth= 13] . ( 29 sec) [ -31585.9074] [ 6374] [depth= 1] . ( 29 sec) [ -31518.8159] [ 6365] [depth= 25] . ( 30 sec) [ -31496.0441] [ 6362] [depth= 23] . ( 30 sec) [ -31453.7469] [ 6366] [depth= 1] . ( 30 sec) [ -30914.4534] [ 6228] [depth= 24] . ( 30 sec) [ -30881.5104] [ 6227] [depth= 1] . ( 31 sec) [ -30808.2225] [ 6214] [depth= 19] . ( 31 sec) [ -30695.8718] [ 6180] [depth= 10] . ( 31 sec) [ -30647.7710] [ 6172] [depth= 1] . ( 31 sec) [ -30606.4657] [ 6167] [depth= 20] . ( 32 sec) [ -30543.7804] [ 6156] [depth= 12] . ( 32 sec) [ -30509.7549] [ 6148] [depth= 10] . ( 32 sec) [ -30482.6106] [ 6150] [depth= 11] . ( 32 sec) [ -30407.1393] [ 6134] [depth= 25] . ( 33 sec) [ -30351.1583] [ 6118] [depth= 20] . ( 33 sec) [ -30277.4620] [ 6092] [depth= 20] . ( 33 sec) [ -30051.7135] [ 6092] [depth= 1] . ( 33 sec) [ -29527.2267] [ 6009] [depth= 20] . ( 34 sec) [ -29409.8232] [ 5952] [depth= 19] . ( 34 sec) [ -29395.6542] [ 5947] [depth= 1] . ( 34 sec) [ -29369.7630] [ 5934] [depth= 1] . ( 35 sec) [ -28651.0884] [ 5925] [depth= 20] . ( 35 sec) [ -28590.2874] [ 5902] [depth= 23] . ( 35 sec) [ -28561.2703] [ 5901] [depth= 1] . ( 35 sec) [ -28554.0850] [ 5895] [depth= 1] . ( 35 sec) [ -28553.9829] [ 5897] [depth= 1] . ( 36 sec) [ -28216.0709] [ 5890] [depth= 8] . ( 36 sec) [ -28209.0107] [ 5888] [depth= 1] . ( 36 sec) [ -28151.8376] [ 5873] [depth= 11] . ( 37 sec) [ -28126.8963] [ 5870] [depth= 27] . ( 37 sec) [ -28114.6555] [ 5863] [depth= 3] . ( 37 sec) [ -28078.5831] [ 5862] [depth= 20] . ( 37 sec) [ -28050.1885] [ 5861] [depth= 2] . ( 38 sec) [ -28019.7279] [ 5862] [depth= 14] . ( 38 sec) [ -27992.0489] [ 5859] [depth= 21] . ( 38 sec) [ -27965.9403] [ 5855] [depth= 5] . ( 38 sec) [ -27961.3412] [ 5856] [depth= 1] . ( 39 sec) [ -27900.4865] [ 5839] [depth= 6] . ( 39 sec) [ -27890.7824] [ 5840] [depth= 18] . ( 39 sec) [ -27880.0340] [ 5833] [depth= 5] . ( 39 sec) [ -27879.2281] [ 5830] [depth= 1] . ( 40 sec) [ -27860.4060] [ 5826] [depth= 4] . ( 40 sec) [ -27717.5211] [ 5763] [depth= 2] . ( 40 sec) [ -27602.7313] [ 5722] [depth= 22] . ( 41 sec) [ -27581.0806] [ 5716] [depth= 1] . ( 41 sec) [ -27579.3892] [ 5718] [depth= 1] . ( 41 sec) [ -27564.8009] [ 5709] [depth= 1] . ( 41 sec) [ -27564.6240] [ 5717] [depth= 1] . ( 42 sec) [ -27534.1173] [ 5710] [depth= 8] . ( 42 sec) [ -27231.0791] [ 5574] [depth= 15] . ( 42 sec) [ -27229.4340] [ 5574] [depth= 1] . ( 42 sec) [ -27014.4229] [ 5573] [depth= 4] . ( 42 sec) [ -26968.5571] [ 5559] [depth= 1] . ( 43 sec) [ -26968.1946] [ 5559] [depth= 1] . ( 43 sec) [ -26960.7356] [ 5558] [depth= 13] . ( 43 sec) [ -26960.0430] [ 5557] [depth= 1] . ( 43 sec) [ -26901.1122] [ 5556] [depth= 12] . ( 44 sec) [ -26875.3630] [ 5550] [depth= 16] . ( 44 sec) [ -26813.9945] [ 5552] [depth= 1] . ( 44 sec) [ -26757.4833] [ 5535] [depth= 11] . ( 44 sec) [ -26749.3846] [ 5531] [depth= 5] . ( 45 sec) [ -26686.6896] [ 5516] [depth= 15] . ( 45 sec) [ -26471.2378] [ 5439] [depth= 15] . ( 45 sec) [ -26467.2589] [ 5446] [depth= 1] . ( 46 sec) [ -26371.2590] [ 5398] [depth= 11] . ( 46 sec) [ -26320.9058] [ 5363] [depth= 12] . ( 46 sec) [ -26307.3721] [ 5368] [depth= 1] . ( 46 sec) [ -26140.7186] [ 5359] [depth= 14] . ( 47 sec) [ -26135.0805] [ 5356] [depth= 1] . ( 47 sec) [ -26081.5884] [ 5354] [depth= 4] . ( 47 sec) [ -25985.8310] [ 5327] [depth= 11] . ( 48 sec) [ -25978.9300] [ 5323] [depth= 3] . ( 48 sec) [ -25948.9536] [ 5315] [depth= 7] . ( 48 sec) [ -25943.7610] [ 5315] [depth= 13] . ( 49 sec) [ -25937.4162] [ 5312] [depth= 11] . ( 49 sec) [ -25932.7634] [ 5311] [depth= 2] . ( 49 sec) [ -25918.5840] [ 5311] [depth= 3] . ( 49 sec) [ -25906.2460] [ 5310] [depth= 3] . ( 50 sec) [ -25885.8622] [ 5306] [depth= 1] . ( 50 sec) [ -25876.7429] [ 5304] [depth= 2] . ( 50 sec) [ -25864.5120] [ 5303] [depth= 5] . ( 50 sec) [ -25771.9147] [ 5281] [depth= 9] . ( 51 sec) [ -25753.6923] [ 5278] [depth= 15] . ( 51 sec) [ -25751.8726] [ 5278] [depth= 1] . ( 51 sec) [ -25689.4315] [ 5262] [depth= 3] . ( 51 sec) [ -25672.5803] [ 5260] [depth= 3] . ( 52 sec) [ -25657.4478] [ 5257] [depth= 2] . ( 52 sec) [ -25628.7113] [ 5257] [depth= 1] . ( 52 sec) [ -25604.2965] [ 5257] [depth= 2] . ( 52 sec) [ -25595.2599] [ 5257] [depth= 3] . ( 53 sec) [ -25571.5403] [ 5252] [depth= 8] . ( 53 sec) [ -25568.7381] [ 5250] [depth= 5] . ( 53 sec) [ -25563.9954] [ 5248] [depth= 11] . ( 54 sec) [ -25514.7548] [ 5220] [depth= 21] . ( 55 sec) [ -25509.7309] [ 5212] [depth= 5] . ( 55 sec) [ -25475.3431] [ 5207] [depth= 2] . ( 55 sec) [ -25367.5057] [ 5139] [depth= 1] . ( 55 sec) [ -25360.8873] [ 5135] [depth= 3] . ( 56 sec) [ -25359.5877] [ 5136] [depth= 1] . ( 56 sec) [ -25356.2127] [ 5135] [depth= 1] . ( 56 sec) [ -25325.6553] [ 5132] [depth= 17] . ( 56 sec) [ -25311.4040] [ 5131] [depth= 1] . ( 57 sec) [ -24977.9899] [ 5059] [depth= 10] . ( 57 sec) [ -24968.3685] [ 5057] [depth= 5] . ( 57 sec) [ -24962.3205] [ 5055] [depth= 1] . ( 59 sec) [ -24961.1493] [ 5058] [depth= 15] . ( 59 sec) [ -24952.0345] [ 5055] [depth= 13] . ( 59 sec) [ -24931.2297] [ 5050] [depth= 2] . ( 59 sec) [ -24927.6121] [ 5049] [depth= 1] . ( 60 sec) [ -24925.9311] [ 5051] [depth= 12] . ( 60 sec) [ -24919.1345] [ 5049] [depth= 1] . ( 60 sec) [ -24909.8471] [ 5048] [depth= 1] . ( 61 sec) [ -24887.0412] [ 5043] [depth= 17] . ( 61 sec) [ -24836.7587] [ 5031] [depth= 11] . ( 61 sec) [ -24833.2287] [ 5032] [depth= 1] . ( 61 sec) [ -24805.0559] [ 5026] [depth= 7] . ( 62 sec) [ -24786.5707] [ 5011] [depth= 22] . ( 62 sec) [ -24785.3124] [ 5011] [depth= 1] . ( 63 sec) [ -24531.1258] [ 4907] [depth= 4] . ( 64 sec) [ -24512.9390] [ 4902] [depth= 8] . ( 64 sec) [ -24492.0727] [ 4897] [depth= 6] . ( 64 sec) [ -24489.8010] [ 4898] [depth= 1] . ( 64 sec) [ -24489.6264] [ 4898] [depth= 1] . ( 65 sec) [ -24481.2801] [ 4895] [depth= 1] . ( 65 sec) [ -24479.6127] [ 4895] [depth= 1] . ( 65 sec) [ -24442.1252] [ 4885] [depth= 1] . ( 65 sec) [ -24420.0105] [ 4876] [depth= 1] . ( 66 sec) [ -24418.8906] [ 4877] [depth= 1] . ( 66 sec) [ -24418.3710] [ 4877] [depth= 2] . ( 66 sec) [ -24407.6896] [ 4875] [depth= 18] . ( 67 sec) [ -24389.7091] [ 4874] [depth= 19] . ( 67 sec) [ -24383.5434] [ 4873] [depth= 6] . ( 67 sec) [ -24380.9729] [ 4870] [depth= 15] . ( 68 sec) [ -24379.0951] [ 4869] [depth= 4] . ( 68 sec) [ -24376.2450] [ 4869] [depth= 1] . ( 68 sec) [ -24371.7937] [ 4866] [depth= 5] . ( 68 sec) [ -24329.3147] [ 4865] [depth= 1] . ( 69 sec) [ -24313.1276] [ 4862] [depth= 2] . ( 69 sec) [ -24182.6101] [ 4856] [depth= 6] . ( 69 sec) [ -24182.3741] [ 4848] [depth= 24] . ( 70 sec) [ -24147.4376] [ 4828] [depth= 26] . ( 70 sec) [ -24138.0064] [ 4821] [depth= 11] . ( 70 sec) [ -24095.6101] [ 4806] [depth= 16] . ( 70 sec) [ -24062.3188] [ 4795] [depth= 1] . ( 71 sec) [ -24038.2645] [ 4851] [depth= 3] . ( 73 sec) [ -24031.8811] [ 4850] [depth= 3] . ( 73 sec) [ -23968.9146] [ 4849] [depth= 1] . ( 73 sec) [ -23761.6445] [ 4825] [depth= 8] . ( 74 sec) [ -23510.9884] [ 4788] [depth= 11] . ( 74 sec) [ -23482.8964] [ 4793] [depth= 13] . ( 74 sec) [ -23416.1880] [ 4767] [depth= 20] . ( 74 sec) [ -23392.0745] [ 4762] [depth= 1] . ( 75 sec) [ -23364.4020] [ 4759] [depth= 5] . ( 75 sec) [ -23329.2091] [ 4769] [depth= 1] . ( 75 sec) [ -23289.7768] [ 4746] [depth= 6] . ( 75 sec) [ -23269.4569] [ 4743] [depth= 2] . ( 76 sec) [ -23266.3444] [ 4742] [depth= 1] . ( 76 sec) [ -23266.3209] [ 4743] [depth= 1] . ( 77 sec) [ -23239.3961] [ 4729] [depth= 1] . ( 77 sec) [ -23201.7995] [ 4718] [depth= 5] . ( 78 sec) [ -23145.3762] [ 4705] [depth= 5] . ( 78 sec) [ -23100.1365] [ 4704] [depth= 1] . ( 78 sec) [ -23096.6489] [ 4704] [depth= 1] . ( 78 sec) [ -23094.5499] [ 4704] [depth= 2] . ( 79 sec) [ -23068.5035] [ 4672] [depth= 10] . ( 79 sec) [ -23053.0618] [ 4674] [depth= 1] . ( 79 sec) [ -23048.1573] [ 4674] [depth= 1] . ( 80 sec) [ -23045.0109] [ 4674] [depth= 1] . ( 80 sec) [ -23042.7489] [ 4672] [depth= 15] . ( 81 sec) [ -23004.8391] [ 4648] [depth= 1] . ( 81 sec) [ -22973.5887] [ 4638] [depth= 10] . ( 81 sec) [ -22969.1518] [ 4636] [depth= 1] . ( 81 sec) [ -22944.2344] [ 4631] [depth= 9] . ( 82 sec) [ -22934.9343] [ 4627] [depth= 1] . ( 82 sec) [ -22931.0856] [ 4625] [depth= 4] . ( 82 sec) [ -22923.0158] [ 4627] [depth= 1] . ( 82 sec) [ -22869.5558] [ 4629] [depth= 1] . ( 83 sec) [ -22833.0622] [ 4610] [depth= 2] . ( 83 sec) [ -22831.7928] [ 4610] [depth= 1] . ( 84 sec) [ -22748.3702] [ 4593] [depth= 17] . ( 84 sec) [ -22698.7768] [ 4582] [depth= 2] . ( 85 sec) [ -22664.3088] [ 4581] [depth= 6] . ( 85 sec) [ -22634.0827] [ 4559] [depth= 21] . ( 85 sec) [ -22606.2280] [ 4561] [depth= 1] . ( 85 sec) [ -22594.2150] [ 4560] [depth= 1] . ( 86 sec) [ -22589.3176] [ 4555] [depth= 9] . ( 86 sec) [ -22565.8650] [ 4554] [depth= 1] . ( 86 sec) [ -22527.0592] [ 4552] [depth= 1] . ( 86 sec) [ -22521.9021] [ 4551] [depth= 1] . ( 86 sec) [ -22519.4823] [ 4551] [depth= 1] . ( 87 sec) [ -22517.5200] [ 4550] [depth= 7] . ( 87 sec) [ -22486.5600] [ 4566] [depth= 1] . ( 87 sec) [ -22446.7733] [ 4550] [depth= 1] . ( 88 sec) [ -22445.2961] [ 4552] [depth= 10] . ( 88 sec) [ -22418.4716] [ 4545] [depth= 4] . ( 88 sec) [ -22391.2216] [ 4541] [depth= 11] . ( 88 sec) [ -22389.0546] [ 4540] [depth= 1] . ( 89 sec) [ -22388.3192] [ 4540] [depth= 1] . ( 89 sec) [ -22370.7790] [ 4535] [depth= 4] . ( 89 sec) [ -22368.9687] [ 4536] [depth= 1] . ( 90 sec) [ -22339.5340] [ 4535] [depth= 13] . ( 90 sec) [ -22329.7629] [ 4532] [depth= 3] . ( 90 sec) [ -22326.8181] [ 4532] [depth= 5] . ( 91 sec) [ -22318.5214] [ 4531] [depth= 4] . ( 91 sec) [ -22306.6402] [ 4528] [depth= 1] . ( 92 sec) [ -22306.0528] [ 4528] [depth= 3] . ( 92 sec) [ -22246.0762] [ 4507] [depth= 6] . ( 92 sec) [ -22245.8224] [ 4507] [depth= 1] . ( 92 sec) [ -22187.7185] [ 4482] [depth= 15] . ( 93 sec) [ -22174.3136] [ 4479] [depth= 1] . ( 93 sec) [ -22149.7916] [ 4475] [depth= 7] . ( 93 sec) [ -22149.7332] [ 4476] [depth= 1] . ( 94 sec) [ -22149.2314] [ 4476] [depth= 1] . ( 94 sec) [ -22137.6476] [ 4474] [depth= 3] . ( 95 sec) [ -22125.3146] [ 4469] [depth= 6] . ( 95 sec) [ -22124.6266] [ 4470] [depth= 1] . ( 96 sec) [ -22097.8207] [ 4439] [depth= 6] . ( 96 sec) [ -22083.3589] [ 4438] [depth= 1] . ( 96 sec) [ -22079.4070] [ 4438] [depth= 1] . ( 96 sec) [ -22078.4982] [ 4438] [depth= 1] . ( 97 sec) [ -22076.8030] [ 4441] [depth= 1] . ( 97 sec) [ -21969.2254] [ 4404] [depth= 2] . ( 97 sec) [ -21887.7120] [ 4404] [depth= 1] . ( 98 sec) [ -21794.8036] [ 4404] [depth= 14] . ( 99 sec) [ -21655.2416] [ 4371] [depth= 19] . ( 99 sec) [ -21599.7192] [ 4353] [depth= 15] . ( 99 sec) [ -21568.3875] [ 4370] [depth= 18] . ( 101 sec) [ -21297.7124] [Branch lengths ] . ( 108 sec) [ -21278.5559] [GTR parameters ] . ( 108 sec) [ -21222.3775] [Alpha ][ 0.345469] . ( 109 sec) [ -21222.3544] [ 4370] [depth= 1] . ( 109 sec) [ -21218.9532] [ 4365] [depth= 2] . ( 109 sec) [ -21217.7999] [ 4366] [depth= 2] . ( 110 sec) [ -21198.0947] [ 4361] [depth= 11] . ( 117 sec) [ -21170.4822] [ 4347] [depth= 11] . ( 118 sec) [ -21169.9731] [ 4347] [depth= 2] . ( 119 sec) [ -21167.8681] [ 4350] [depth= 1] . ( 119 sec) [ -21166.2269] [ 4346] [depth= 13] . ( 119 sec) [ -21160.6607] [ 4347] [depth= 2] . ( 119 sec) [ -21160.0386] [ 4348] [depth= 1] . ( 120 sec) [ -21156.4684] [ 4347] [depth= 1] . ( 123 sec) [ -21155.6377] [ 4343] [depth= 1] . ( 123 sec) [ -21144.0429] [ 4339] [depth= 1] . ( 125 sec) [ -21141.1707] [ 4345] [depth= 5] . ( 125 sec) [ -21116.7547] [ 4330] [depth= 5] . ( 126 sec) [ -21115.2758] [ 4327] [depth= 1] . ( 127 sec) [ -21113.3794] [ 4328] [depth= 1] . ( 127 sec) [ -21044.2579] [ 4297] [depth= 15] . ( 127 sec) [ -21044.2477] [ 4297] [depth= 1] . ( 128 sec) [ -21043.9341] [ 4297] [depth= 1] . ( 128 sec) [ -21043.4797] [ 4296] [depth= 1] . ( 128 sec) [ -21043.0042] [ 4296] [depth= 1] . ( 129 sec) [ -21042.7773] [ 4292] [depth= 3] . ( 129 sec) [ -21042.7474] [ 4292] [depth= 1] . ( 130 sec) [ -21023.7529] [ 4271] [depth= 2] . ( 130 sec) [ -21021.1742] [ 4265] [depth= 12] . ( 133 sec) [ -21010.0452] [ 4263] [depth= 1] . ( 137 sec) [ -21009.4266] [ 4264] [depth= 1] . ( 137 sec) [ -21007.5284] [ 4264] [depth= 2] . ( 137 sec) [ -20971.3588] [ 4253] [depth= 1] . ( 138 sec) [ -20905.7702] [ 4228] [depth= 12] . ( 140 sec) [ -20899.7227] [ 4222] [depth= 2] . ( 143 sec) [ -20899.4360] [ 4222] [depth= 1] . ( 143 sec) [ -20897.9727] [ 4222] [depth= 1] . ( 144 sec) [ -20897.8905] [ 4222] [depth= 2] . ( 144 sec) [ -20896.7030] [ 4220] [depth= 2] . ( 146 sec) [ -20895.5962] [ 4220] [depth= 3] . ( 147 sec) [ -20895.2335] [ 4219] [depth= 2] . ( 150 sec) [ -20895.0011] [ 4217] [depth= 1] . ( 150 sec) [ -20894.9162] [ 4218] [depth= 1] . ( 154 sec) [ -20894.8392] [ 4218] [depth= 1] . ( 155 sec) [ -20885.1365] [ 4206] [depth= 9] . ( 155 sec) [ -20849.1711] [ 4191] [depth= 1] . ( 157 sec) [ -20848.3908] [ 4191] [depth= 3] . ( 157 sec) [ -20845.1738] [ 4190] [depth= 1] . ( 158 sec) [ -20844.2368] [ 4190] [depth= 1] . ( 161 sec) [ -20832.6508] [ 4185] [depth= 1] . ( 162 sec) [ -20831.5552] [ 4187] [depth= 1] . ( 163 sec) [ -20818.1340] [ 4184] [depth= 3] . ( 163 sec) [ -20818.0821] [ 4185] [depth= 2] . ( 165 sec) [ -20817.7519] [ 4187] [depth= 1] . ( 166 sec) [ -20814.8980] [ 4186] [depth= 1] . ( 170 sec) [ -20773.6829] [ 4165] [depth= 2] . ( 171 sec) [ -20770.8081] [ 4168] [depth= 1] . ( 173 sec) [ -20759.3449] [ 4164] [depth= 2] . ( 173 sec) [ -20758.0745] [ 4165] [depth= 1] . ( 174 sec) [ -20758.0258] [ 4165] [depth= 1] . ( 175 sec) [ -20756.4970] [ 4164] [depth= 2] . ( 177 sec) [ -20755.6637] [ 4164] [depth= 1] . ( 178 sec) [ -20755.1674] [ 4164] [depth= 1] . ( 178 sec) [ -20753.1932] [ 4158] [depth= 7] . ( 181 sec) [ -20741.2608] [ 4152] [depth= 1] . ( 182 sec) [ -20709.9238] [ 4137] [depth= 2] . ( 184 sec) [ -20708.8588] [ 4135] [depth= 2] . ( 184 sec) [ -20705.0882] [ 4136] [depth= 1] . ( 185 sec) [ -20702.4763] [ 4137] [depth= 1] . ( 185 sec) [ -20698.5095] [ 4135] [depth= 2] . ( 185 sec) [ -20680.7694] [ 4105] [depth= 1] . ( 185 sec) [ -20680.4830] [ 4105] [depth= 1] . ( 186 sec) [ -20679.1070] [ 4104] [depth= 4] . ( 187 sec) [ -20678.5472] [ 4104] [depth= 2] . ( 188 sec) [ -20678.2998] [ 4104] [depth= 1] . ( 188 sec) [ -20675.0883] [ 4106] [depth= 2] . ( 189 sec) [ -20674.8243] [ 4106] [depth= 1] . ( 190 sec) [ -20674.4309] [ 4104] [depth= 2] . ( 190 sec) [ -20674.3462] [ 4104] [depth= 1] . ( 193 sec) [ -20672.7956] [ 4103] [depth= 2] . ( 193 sec) [ -20667.3861] [ 4107] [depth= 10] . ( 193 sec) [ -20666.6364] [ 4098] [depth= 1] . ( 194 sec) [ -20666.4826] [ 4098] [depth= 1] . ( 195 sec) [ -20666.4486] [ 4098] [depth= 1] . ( 196 sec) [ -20661.8546] [ 4098] [depth= 2] . ( 197 sec) [ -20660.1844] [ 4098] [depth= 1] . ( 198 sec) [ -20659.6448] [ 4098] [depth= 1] . ( 199 sec) [ -20657.4338] [ 4096] [depth= 3] . ( 199 sec) [ -20654.1781] [ 4097] [depth= 17] . ( 200 sec) [ -20645.9881] [ 4092] [depth= 1] . ( 200 sec) [ -20645.8730] [ 4093] [depth= 1] . ( 201 sec) [ -20638.9772] [ 4094] [depth= 1] . ( 201 sec) [ -20637.9075] [ 4096] [depth= 1] . ( 201 sec) [ -20637.6526] [ 4099] [depth= 1] . ( 202 sec) [ -20637.5458] [ 4099] [depth= 1] . ( 202 sec) [ -20636.7051] [ 4099] [depth= 2] . ( 202 sec) [ -20636.0996] [ 4098] [depth= 2] . ( 202 sec) [ -20629.9184] [ 4098] [depth= 2] . ( 203 sec) [ -20629.8128] [ 4101] [depth= 5] . ( 205 sec) [ -20621.7963] [ 4098] [depth= 1] . ( 207 sec) [ -20620.7590] [ 4102] [depth= 2] . ( 207 sec) [ -20619.1636] [ 4101] [depth= 1] . ( 208 sec) [ -20617.8878] [ 4101] [depth= 1] . ( 208 sec) [ -20609.2935] [ 4105] [depth= 1] . ( 211 sec) [ -20599.5482] [ 4103] [depth= 2] . ( 211 sec) [ -20596.4382] [ 4102] [depth= 1] . ( 213 sec) [ -20596.0714] [ 4101] [depth= 1] . ( 213 sec) [ -20594.9682] [ 4102] [depth= 1] . ( 213 sec) [ -20594.8221] [ 4102] [depth= 1] . ( 214 sec) [ -20594.6815] [ 4103] [depth= 1] . ( 215 sec) [ -20594.3958] [ 4103] [depth= 1] . ( 215 sec) [ -20594.1781] [ 4104] [depth= 1] . ( 215 sec) [ -20576.7979] [ 4097] [depth= 10] . ( 216 sec) [ -20576.2484] [ 4096] [depth= 1] . ( 216 sec) [ -20573.5660] [ 4094] [depth= 1] . ( 217 sec) [ -20562.6176] [ 4093] [depth= 2] . ( 218 sec) [ -20562.2535] [ 4093] [depth= 1] . ( 219 sec) [ -20561.7033] [ 4091] [depth= 1] . ( 219 sec) [ -20561.4842] [ 4095] [depth= 1] . ( 220 sec) [ -20559.3822] [ 4095] [depth= 3] . ( 222 sec) [ -20551.0518] [ 4091] [depth= 3] . ( 222 sec) [ -20549.8497] [ 4093] [depth= 3] . ( 223 sec) [ -20540.5688] [ 4093] [depth= 1] . ( 223 sec) [ -20540.3078] [ 4090] [depth= 1] . ( 226 sec) [ -20540.2139] [ 4090] [depth= 1] . ( 227 sec) [ -20537.8379] [ 4088] [depth= 2] . ( 227 sec) [ -20536.9690] [ 4079] [depth= 11] . ( 229 sec) [ -20536.8763] [ 4079] [depth= 1] . ( 229 sec) [ -20530.4967] [ 4077] [depth= 6] . ( 230 sec) [ -20528.8894] [ 4075] [depth= 1] . ( 233 sec) [ -20462.4836] [Branch lengths ] . ( 236 sec) [ -20461.1778] [GTR parameters ] . ( 237 sec) [ -20461.0561] [Alpha ][ 0.339903] . ( 238 sec) [ -20459.4734] [ 4070] [depth= 5] . ( 244 sec) [ -20458.5919] [ 4069] [depth= 1] . ( 249 sec) [ -20456.0630] [ 4069] [depth= 1] . ( 251 sec) [ -20453.5055] [ 4067] [depth= 1] . ( 255 sec) [ -20453.4020] [ 4067] [depth= 1] . ( 260 sec) [ -20452.9307] [ 4065] [depth= 1] . ( 263 sec) [ -20452.3467] [ 4064] [depth= 1] . ( 272 sec) [ -20451.2351] [ 4060] [depth= 3] . ( 274 sec) [ -20451.0575] [ 4061] [depth= 1] . ( 274 sec) [ -20450.2232] [ 4060] [depth= 3] . ( 275 sec) [ -20449.9098] [ 4060] [depth= 1] . ( 276 sec) [ -20448.2690] [ 4059] [depth= 1] . ( 276 sec) [ -20447.2117] [ 4059] [depth= 3] . ( 278 sec) [ -20446.6101] [ 4057] [depth= 1] . ( 282 sec) [ -20445.7590] [ 4058] [depth= 3] . ( 284 sec) [ -20445.5439] [ 4062] [depth= 1] . ( 284 sec) [ -20445.4688] [ 4063] [depth= 1] . ( 291 sec) [ -20444.1411] [ 4062] [depth= 1] . ( 297 sec) [ -20443.5274] [ 4064] [depth= 1] . ( 298 sec) [ -20443.3584] [ 4064] [depth= 1] . ( 300 sec) [ -20442.9416] [ 4064] [depth= 1] . ( 301 sec) [ -20440.6046] [ 4057] [depth= 1] . ( 302 sec) [ -20440.4372] [ 4057] [depth= 1] . ( 302 sec) [ -20439.4731] [ 4057] [depth= 1] . ( 303 sec) [ -20438.5365] [ 4057] [depth= 1] . ( 315 sec) [ -20436.0869] [ 4056] [depth= 2] . ( 316 sec) [ -20435.8649] [ 4056] [depth= 1] . ( 317 sec) [ -20435.2353] [ 4055] [depth= 1] . ( 319 sec) [ -20435.1435] [ 4055] [depth= 1] . ( 322 sec) [ -20434.0425] [ 4056] [depth= 1] . ( 326 sec) [ -20433.7608] [ 4056] [depth= 2] . ( 334 sec) [ -20433.2789] [ 4061] [depth= 1] . ( 336 sec) [ -20433.1595] [ 4061] [depth= 1] . ( 338 sec) [ -20433.1031] [ 4060] [depth= 1] . ( 339 sec) [ -20433.0078] [ 4060] [depth= 1] . ( 340 sec) [ -20432.7133] [ 4060] [depth= 1] . ( 345 sec) [ -20432.1316] [ 4060] [depth= 1] . ( 349 sec) [ -20430.8664] [ 4059] [depth= 1] . ( 353 sec) [ -20429.7586] [ 4053] [depth= 4] . ( 360 sec) [ -20416.2299] [Branch lengths ] . ( 363 sec) [ -20416.1549] [GTR parameters ] . ( 363 sec) [ -20416.1499] [Alpha ][ 0.338640] . ( 366 sec) [ -20415.3006] [ 4054] [depth= 1] . ( 463 sec) [ -20415.1503] [ 4058] [depth= 9] . ( 480 sec) [ -20415.0381] [ 4061] [depth= 1] . ( 491 sec) [ -20414.9657] [ 4061] [depth= 2] . ( 503 sec) [ -20414.9039] [ 4060] [depth= 1] . ( 509 sec) [ -20411.3292] [Branch lengths ] . ( 512 sec) [ -20411.2862] [GTR parameters ] . ( 513 sec) [ -20411.2850] [Alpha ][ 0.339121] . ( 514 sec) [ -20409.2448] [ 4062] [depth= 2] . ( 538 sec) [ -20409.1776] [ 4058] [depth= 1] . ( 540 sec) [ -20408.4376] [ 4057] [depth= 1] . ( 563 sec) [ -20407.8936] [ 4055] [depth= 1] . ( 563 sec) [ -20407.2730] [ 4053] [depth= 9] . ( 598 sec) [ -20406.6456] [ 4054] [depth= 8] . ( 600 sec) [ -20405.4348] [ 4053] [depth= 1] . ( 611 sec) [ -20404.1382] [ 4053] [depth= 1] . ( 613 sec) [ -20404.1240] [ 4053] [depth= 1] . ( 641 sec) [ -20404.0455] [ 4053] [depth= 2] . ( 644 sec) [ -20398.5951] [Branch lengths ] . ( 647 sec) [ -20398.5534] [GTR parameters ] . ( 648 sec) [ -20398.5523] [Alpha ][ 0.339583] . ( 737 sec) [ -20398.5379] [ 4053] [depth= 1] . ( 778 sec) [ -20398.4061] [Branch lengths ] . ( 781 sec) [ -20398.4054] [GTR parameters ] . ( 781 sec) [ -20398.4050] [Alpha ][ 0.339923] . ( 1020 sec) [ -20397.7196] [ 4066] [depth= 2] . ( 1027 sec) [ -20397.2674] [ 4065] [depth= 2] . ( 1032 sec) [ -20397.1877] [Branch lengths ] . ( 1036 sec) [ -20397.1837] [GTR parameters ] . ( 1036 sec) [ -20397.1837] [Alpha ][ 0.339923] . ( 1117 sec) [ -20396.9176] [ 4064] [depth= 1] . ( 1150 sec) [ -20396.4878] [ 4065] [depth= 1] . ( 1268 sec) [ -20396.3624] [ 4064] [depth= 1] . ( 1286 sec) [ -20395.8225] [Branch lengths ] . ( 1290 sec) [ -20395.8211] [GTR parameters ] . ( 1290 sec) [ -20395.8205] [Alpha ][ 0.340263] . ( 1542 sec) [ -20395.7960] [Branch lengths ] . ( 1542 sec) [ -20395.7960] [Topology ] . ( 1547 sec) [ -20395.7920] [Topology ] . ( 1552 sec) [ -20395.7899] [Topology ] . ( 1557 sec) [ -20395.7886] [Topology ] . ( 1563 sec) [ -20395.7877] [Topology ] . ( 1563 sec) [ -20395.7877] [GTR parameters ] . ( 1564 sec) [ -20395.7877] [Alpha ][ 0.340263] . ( 1564 sec) [ -20395.7877] [Topology ] . ( 1569 sec) [ -20395.7870] [Topology ] . ( 1570 sec) [ -20395.7870] [GTR parameters ] . ( 1570 sec) [ -20395.7870] [Alpha ][ 0.340263] . Checking for NNIs, optimizing five branches... . ( 1608 sec) [ -20395.7336] [Topology ] . Checking for NNIs, optimizing five branches... . Log likelihood of the current tree: -20395.733581. . [Random start 5/ 5] . Computing pairwise distances... . Building BioNJ tree... . Randomising the tree... . Maximizing likelihood (using SPR moves)... . ( 5 sec) [ -45647.5279] [GTR parameters ] . ( 6 sec) [ -44575.6309] [Alpha ][ 0.500960] . ( 7 sec) [ -44496.1342] [ 7563] [depth= 12] . ( 7 sec) [ -44426.7579] [ 7551] [depth= 8] . ( 7 sec) [ -44322.3023] [ 7550] [depth= 1] . ( 7 sec) [ -44315.9363] [ 7547] [depth= 7] . ( 8 sec) [ -44313.4923] [ 7545] [depth= 2] . ( 8 sec) [ -44310.5954] [ 7539] [depth= 15] . ( 8 sec) [ -44300.9673] [ 7537] [depth= 4] . ( 8 sec) [ -44280.2348] [ 7531] [depth= 6] . ( 9 sec) [ -44278.5384] [ 7526] [depth= 10] . ( 9 sec) [ -44212.2245] [ 7525] [depth= 1] . ( 9 sec) [ -44142.0200] [ 7524] [depth= 3] . ( 9 sec) [ -44104.7918] [ 7524] [depth= 3] . ( 9 sec) [ -44095.0528] [ 7522] [depth= 1] . ( 10 sec) [ -44075.4180] [ 7520] [depth= 4] . ( 10 sec) [ -44059.1270] [ 7520] [depth= 5] . ( 10 sec) [ -43952.0258] [ 7519] [depth= 3] . ( 10 sec) [ -43847.8189] [ 7492] [depth= 4] . ( 11 sec) [ -43692.1419] [ 7461] [depth= 8] . ( 11 sec) [ -43637.7570] [ 7461] [depth= 2] . ( 11 sec) [ -43528.2622] [ 7456] [depth= 8] . ( 11 sec) [ -43517.9403] [ 7456] [depth= 2] . ( 11 sec) [ -43516.0743] [ 7456] [depth= 1] . ( 12 sec) [ -43515.7333] [ 7456] [depth= 1] . ( 12 sec) [ -43468.0977] [ 7450] [depth= 9] . ( 12 sec) [ -43434.2244] [ 7442] [depth= 3] . ( 13 sec) [ -43367.2708] [ 7424] [depth= 35] . ( 13 sec) [ -43337.4424] [ 7415] [depth= 22] . ( 13 sec) [ -42889.7794] [ 7390] [depth= 34] . ( 13 sec) [ -42587.7503] [ 7296] [depth= 14] . ( 14 sec) [ -42396.2294] [ 7318] [depth= 31] . ( 14 sec) [ -42249.8355] [ 7273] [depth= 26] . ( 14 sec) [ -42249.6705] [ 7277] [depth= 1] . ( 14 sec) [ -41778.2847] [ 7268] [depth= 1] . ( 15 sec) [ -41777.3629] [ 7268] [depth= 2] . ( 15 sec) [ -41727.1381] [ 7267] [depth= 3] . ( 15 sec) [ -41725.0832] [ 7265] [depth= 5] . ( 15 sec) [ -41703.9231] [ 7263] [depth= 1] . ( 15 sec) [ -41670.0874] [ 7256] [depth= 9] . ( 16 sec) [ -41184.3262] [ 7203] [depth= 8] . ( 16 sec) [ -41009.5992] [ 7199] [depth= 16] . ( 16 sec) [ -40979.4808] [ 7193] [depth= 6] . ( 17 sec) [ -40975.0603] [ 7193] [depth= 3] . ( 17 sec) [ -40942.9421] [ 7192] [depth= 1] . ( 17 sec) [ -40622.7040] [ 7158] [depth= 16] . ( 17 sec) [ -40479.4326] [ 7156] [depth= 1] . ( 18 sec) [ -40192.2415] [ 7098] [depth= 10] . ( 18 sec) [ -40172.8560] [ 7099] [depth= 5] . ( 18 sec) [ -39770.5377] [ 7036] [depth= 8] . ( 18 sec) [ -39683.5809] [ 7015] [depth= 9] . ( 19 sec) [ -39586.7382] [ 7000] [depth= 18] . ( 19 sec) [ -39089.1725] [ 6982] [depth= 5] . ( 19 sec) [ -39001.4639] [ 6972] [depth= 28] . ( 20 sec) [ -38750.3916] [ 6864] [depth= 20] . ( 20 sec) [ -38469.7138] [ 6858] [depth= 8] . ( 20 sec) [ -38455.4548] [ 6855] [depth= 3] . ( 20 sec) [ -38180.9497] [ 6854] [depth= 2] . ( 21 sec) [ -37940.2211] [ 6851] [depth= 8] . ( 21 sec) [ -37919.1865] [ 6851] [depth= 1] . ( 21 sec) [ -37510.4388] [ 6832] [depth= 11] . ( 21 sec) [ -37468.7607] [ 6830] [depth= 1] . ( 22 sec) [ -37427.3659] [ 6833] [depth= 2] . ( 22 sec) [ -37426.9959] [ 6832] [depth= 4] . ( 22 sec) [ -37414.0103] [ 6831] [depth= 1] . ( 22 sec) [ -37401.0794] [ 6828] [depth= 2] . ( 23 sec) [ -37334.8766] [ 6829] [depth= 7] . ( 23 sec) [ -37319.7366] [ 6827] [depth= 3] . ( 23 sec) [ -37282.4489] [ 6819] [depth= 25] . ( 23 sec) [ -37127.1437] [ 6813] [depth= 18] . ( 24 sec) [ -36874.2082] [ 6682] [depth= 1] . ( 24 sec) [ -36862.6735] [ 6678] [depth= 1] . ( 24 sec) [ -36732.3060] [ 6646] [depth= 8] . ( 25 sec) [ -36668.2902] [ 6620] [depth= 14] . ( 25 sec) [ -36612.1543] [ 6615] [depth= 7] . ( 25 sec) [ -36508.3716] [ 6606] [depth= 12] . ( 25 sec) [ -36502.1936] [ 6607] [depth= 1] . ( 26 sec) [ -36473.7903] [ 6604] [depth= 6] . ( 26 sec) [ -36455.7823] [ 6599] [depth= 21] . ( 26 sec) [ -36436.7924] [ 6595] [depth= 27] . ( 26 sec) [ -36396.9481] [ 6593] [depth= 2] . ( 27 sec) [ -36343.7939] [ 6585] [depth= 11] . ( 27 sec) [ -36330.0845] [ 6586] [depth= 1] . ( 27 sec) [ -36313.9611] [ 6582] [depth= 1] . ( 27 sec) [ -36087.9497] [ 6583] [depth= 1] . ( 27 sec) [ -35984.4908] [ 6562] [depth= 7] . ( 28 sec) [ -35959.7293] [ 6557] [depth= 7] . ( 28 sec) [ -35946.7845] [ 6557] [depth= 1] . ( 28 sec) [ -35733.0712] [ 6484] [depth= 10] . ( 28 sec) [ -35677.3234] [ 6469] [depth= 14] . ( 29 sec) [ -35675.5974] [ 6469] [depth= 1] . ( 29 sec) [ -35651.3151] [ 6464] [depth= 8] . ( 29 sec) [ -35641.2043] [ 6460] [depth= 5] . ( 29 sec) [ -35632.9038] [ 6460] [depth= 1] . ( 30 sec) [ -35554.4566] [ 6458] [depth= 15] . ( 30 sec) [ -35534.3060] [ 6459] [depth= 8] . ( 30 sec) [ -35490.3156] [ 6458] [depth= 1] . ( 30 sec) [ -35454.9925] [ 6452] [depth= 6] . ( 31 sec) [ -34731.9325] [ 6424] [depth= 22] . ( 31 sec) [ -34514.1682] [ 6420] [depth= 1] . ( 31 sec) [ -34315.5399] [ 6379] [depth= 17] . ( 31 sec) [ -33927.6145] [ 6371] [depth= 13] . ( 32 sec) [ -33821.3773] [ 6310] [depth= 10] . ( 32 sec) [ -32267.0318] [ 6316] [depth= 24] . ( 32 sec) [ -32058.3990] [ 6270] [depth= 1] . ( 33 sec) [ -31894.8313] [ 6238] [depth= 19] . ( 33 sec) [ -31883.1942] [ 6231] [depth= 6] . ( 33 sec) [ -31816.3355] [ 6229] [depth= 23] . ( 33 sec) [ -31780.3959] [ 6182] [depth= 14] . ( 33 sec) [ -31749.2458] [ 6176] [depth= 1] . ( 34 sec) [ -31686.0383] [ 6169] [depth= 13] . ( 34 sec) [ -31624.2438] [ 6149] [depth= 31] . ( 34 sec) [ -31589.5797] [ 6140] [depth= 1] . ( 34 sec) [ -31549.9679] [ 6135] [depth= 16] . ( 35 sec) [ -31379.3376] [ 6114] [depth= 21] . ( 35 sec) [ -31285.7830] [ 6086] [depth= 1] . ( 35 sec) [ -31076.9634] [ 6077] [depth= 7] . ( 36 sec) [ -30771.5002] [ 6032] [depth= 1] . ( 36 sec) [ -30540.4531] [ 6009] [depth= 17] . ( 36 sec) [ -30471.3966] [ 5996] [depth= 21] . ( 36 sec) [ -30417.9935] [ 5982] [depth= 2] . ( 37 sec) [ -30416.8664] [ 5986] [depth= 1] . ( 37 sec) [ -30358.0051] [ 5985] [depth= 12] . ( 37 sec) [ -30342.1455] [ 5983] [depth= 8] . ( 38 sec) [ -30329.7598] [ 5982] [depth= 1] . ( 38 sec) [ -30142.6220] [ 5948] [depth= 6] . ( 38 sec) [ -30139.8796] [ 5945] [depth= 2] . ( 38 sec) [ -30136.5137] [ 5931] [depth= 12] . ( 39 sec) [ -30084.0784] [ 5894] [depth= 19] . ( 39 sec) [ -29991.2951] [ 5878] [depth= 14] . ( 39 sec) [ -29969.4176] [ 5865] [depth= 16] . ( 39 sec) [ -29952.3126] [ 5850] [depth= 1] . ( 40 sec) [ -29906.0826] [ 5787] [depth= 1] . ( 40 sec) [ -29842.7692] [ 5736] [depth= 15] . ( 40 sec) [ -29725.7992] [ 5713] [depth= 26] . ( 40 sec) [ -29669.3479] [ 5702] [depth= 1] . ( 41 sec) [ -29617.0897] [ 5691] [depth= 15] . ( 41 sec) [ -29616.9491] [ 5744] [depth= 17] . ( 41 sec) [ -29551.0346] [ 5696] [depth= 7] . ( 41 sec) [ -29548.6762] [ 5696] [depth= 1] . ( 42 sec) [ -29535.3688] [ 5696] [depth= 8] . ( 42 sec) [ -29531.7134] [ 5695] [depth= 1] . ( 42 sec) [ -29501.3285] [ 5689] [depth= 10] . ( 42 sec) [ -29383.2479] [ 5632] [depth= 1] . ( 43 sec) [ -29091.1678] [ 5606] [depth= 15] . ( 43 sec) [ -29066.4257] [ 5599] [depth= 2] . ( 43 sec) [ -29058.1887] [ 5593] [depth= 19] . ( 44 sec) [ -29038.4486] [ 5592] [depth= 3] . ( 44 sec) [ -28998.9734] [ 5593] [depth= 1] . ( 44 sec) [ -28971.4397] [ 5596] [depth= 4] . ( 44 sec) [ -28917.4536] [ 5549] [depth= 4] . ( 45 sec) [ -28909.7206] [ 5551] [depth= 15] . ( 45 sec) [ -28862.3335] [ 5539] [depth= 22] . ( 45 sec) [ -28750.8427] [ 5516] [depth= 11] . ( 45 sec) [ -28745.2422] [ 5515] [depth= 1] . ( 46 sec) [ -28737.1480] [ 5510] [depth= 4] . ( 46 sec) [ -28725.1541] [ 5509] [depth= 1] . ( 46 sec) [ -28699.5860] [ 5512] [depth= 1] . ( 46 sec) [ -28581.4605] [ 5463] [depth= 1] . ( 46 sec) [ -28292.9945] [ 5438] [depth= 9] . ( 47 sec) [ -28284.9257] [ 5433] [depth= 7] . ( 47 sec) [ -28265.5265] [ 5426] [depth= 1] . ( 47 sec) [ -28255.8393] [ 5418] [depth= 6] . ( 48 sec) [ -28206.6959] [ 5401] [depth= 14] . ( 48 sec) [ -28184.3136] [ 5402] [depth= 6] . ( 48 sec) [ -28158.4017] [ 5397] [depth= 1] . ( 48 sec) [ -27571.9673] [ 5367] [depth= 13] . ( 48 sec) [ -27485.8110] [ 5355] [depth= 1] . ( 49 sec) [ -27485.5966] [ 5355] [depth= 1] . ( 49 sec) [ -27424.5099] [ 5354] [depth= 5] . ( 49 sec) [ -27345.1004] [ 5336] [depth= 15] . ( 50 sec) [ -27219.7925] [ 5306] [depth= 9] . ( 50 sec) [ -27213.4415] [ 5300] [depth= 10] . ( 50 sec) [ -27212.2761] [ 5299] [depth= 1] . ( 51 sec) [ -27165.8469] [ 5279] [depth= 11] . ( 51 sec) [ -27082.3174] [ 5277] [depth= 4] . ( 51 sec) [ -27074.9469] [ 5276] [depth= 1] . ( 51 sec) [ -27074.1905] [ 5278] [depth= 2] . ( 52 sec) [ -27044.2005] [ 5273] [depth= 2] . ( 52 sec) [ -27024.5426] [ 5270] [depth= 2] . ( 52 sec) [ -27018.9112] [ 5270] [depth= 1] . ( 52 sec) [ -27012.7100] [ 5267] [depth= 1] . ( 53 sec) [ -26996.5460] [ 5263] [depth= 6] . ( 54 sec) [ -26989.2203] [ 5267] [depth= 7] . ( 54 sec) [ -26987.8393] [ 5267] [depth= 8] . ( 54 sec) [ -26980.4900] [ 5264] [depth= 1] . ( 55 sec) [ -26976.0014] [ 5265] [depth= 10] . ( 55 sec) [ -26906.3716] [ 5249] [depth= 12] . ( 55 sec) [ -26903.9308] [ 5249] [depth= 1] . ( 56 sec) [ -26902.7808] [ 5249] [depth= 1] . ( 56 sec) [ -26751.1889] [ 5180] [depth= 10] . ( 56 sec) [ -26749.4470] [ 5179] [depth= 1] . ( 57 sec) [ -26727.1752] [ 5172] [depth= 5] . ( 57 sec) [ -26713.6331] [ 5164] [depth= 16] . ( 57 sec) [ -26713.4823] [ 5165] [depth= 1] . ( 57 sec) [ -26711.7403] [ 5162] [depth= 1] . ( 58 sec) [ -26695.2208] [ 5154] [depth= 16] . ( 58 sec) [ -26659.7014] [ 5151] [depth= 2] . ( 58 sec) [ -26636.1697] [ 5144] [depth= 1] . ( 58 sec) [ -26574.6554] [ 5139] [depth= 8] . ( 59 sec) [ -26557.7934] [ 5137] [depth= 7] . ( 59 sec) [ -26451.4966] [ 5087] [depth= 3] . ( 59 sec) [ -26445.0446] [ 5085] [depth= 6] . ( 60 sec) [ -26441.4200] [ 5086] [depth= 1] . ( 60 sec) [ -26417.1539] [ 5081] [depth= 2] . ( 60 sec) [ -26393.5569] [ 5077] [depth= 11] . ( 61 sec) [ -26391.2339] [ 5076] [depth= 1] . ( 61 sec) [ -26390.6722] [ 5076] [depth= 1] . ( 62 sec) [ -26372.7274] [ 5074] [depth= 3] . ( 62 sec) [ -26365.7349] [ 5077] [depth= 1] . ( 62 sec) [ -26317.5347] [ 5070] [depth= 9] . ( 62 sec) [ -26229.7485] [ 5068] [depth= 1] . ( 63 sec) [ -26175.3970] [ 5062] [depth= 7] . ( 63 sec) [ -26096.2551] [ 5049] [depth= 15] . ( 63 sec) [ -25960.5397] [ 5039] [depth= 3] . ( 64 sec) [ -25937.3221] [ 5049] [depth= 8] . ( 64 sec) [ -25917.4470] [ 5052] [depth= 2] . ( 64 sec) [ -25847.5417] [ 5030] [depth= 12] . ( 64 sec) [ -25795.9692] [ 5019] [depth= 2] . ( 65 sec) [ -25553.0525] [ 5005] [depth= 12] . ( 65 sec) [ -25530.7362] [ 4979] [depth= 17] . ( 65 sec) [ -25525.9812] [ 4976] [depth= 6] . ( 65 sec) [ -25489.9883] [ 4972] [depth= 3] . ( 66 sec) [ -25479.1488] [ 4971] [depth= 1] . ( 66 sec) [ -25388.4708] [ 4971] [depth= 4] . ( 66 sec) [ -25386.3740] [ 4970] [depth= 11] . ( 66 sec) [ -25349.7382] [ 4970] [depth= 3] . ( 66 sec) [ -25344.6754] [ 4968] [depth= 1] . ( 67 sec) [ -25332.7243] [ 4965] [depth= 1] . ( 67 sec) [ -25316.0711] [ 4965] [depth= 14] . ( 67 sec) [ -25122.7817] [ 4897] [depth= 1] . ( 67 sec) [ -25062.5063] [ 4867] [depth= 3] . ( 68 sec) [ -25044.5368] [ 4865] [depth= 1] . ( 68 sec) [ -25038.8302] [ 4863] [depth= 15] . ( 68 sec) [ -24280.0605] [ 4867] [depth= 13] . ( 68 sec) [ -24242.2115] [ 4843] [depth= 3] . ( 68 sec) [ -24212.8430] [ 4826] [depth= 13] . ( 69 sec) [ -24162.2453] [ 4822] [depth= 1] . ( 70 sec) [ -24148.4615] [ 4822] [depth= 11] . ( 70 sec) [ -24145.3997] [ 4822] [depth= 1] . ( 70 sec) [ -24132.4828] [ 4820] [depth= 8] . ( 70 sec) [ -24123.8812] [ 4819] [depth= 1] . ( 71 sec) [ -24116.3526] [ 4817] [depth= 5] . ( 71 sec) [ -24110.8090] [ 4814] [depth= 10] . ( 71 sec) [ -24069.0431] [ 4811] [depth= 10] . ( 72 sec) [ -24066.8469] [ 4812] [depth= 2] . ( 72 sec) [ -23712.2648] [ 4816] [depth= 6] . ( 72 sec) [ -23663.4635] [ 4804] [depth= 4] . ( 73 sec) [ -23658.0204] [ 4809] [depth= 14] . ( 73 sec) [ -23652.5044] [ 4808] [depth= 1] . ( 73 sec) [ -23650.4828] [ 4807] [depth= 1] . ( 73 sec) [ -23590.9097] [ 4792] [depth= 4] . ( 74 sec) [ -23588.0077] [ 4792] [depth= 1] . ( 74 sec) [ -23587.0809] [ 4791] [depth= 2] . ( 74 sec) [ -23567.2950] [ 4786] [depth= 9] . ( 75 sec) [ -23567.0394] [ 4785] [depth= 2] . ( 75 sec) [ -23560.0260] [ 4786] [depth= 1] . ( 75 sec) [ -23558.1062] [ 4786] [depth= 1] . ( 75 sec) [ -23554.6112] [ 4784] [depth= 3] . ( 76 sec) [ -23535.4898] [ 4787] [depth= 10] . ( 76 sec) [ -23531.5160] [ 4786] [depth= 2] . ( 76 sec) [ -23522.5319] [ 4784] [depth= 4] . ( 76 sec) [ -23514.7769] [ 4779] [depth= 3] . ( 77 sec) [ -23414.3815] [ 4797] [depth= 2] . ( 77 sec) [ -23345.4944] [ 4784] [depth= 14] . ( 77 sec) [ -23283.6706] [ 4733] [depth= 4] . ( 78 sec) [ -23276.6815] [ 4725] [depth= 1] . ( 78 sec) [ -23263.9799] [ 4726] [depth= 3] . ( 78 sec) [ -23262.5464] [ 4725] [depth= 6] . ( 78 sec) [ -23174.1968] [ 4721] [depth= 4] . ( 79 sec) [ -23119.5243] [ 4704] [depth= 6] . ( 79 sec) [ -23117.0060] [ 4704] [depth= 1] . ( 79 sec) [ -23115.9837] [ 4699] [depth= 2] . ( 79 sec) [ -23111.1212] [ 4698] [depth= 1] . ( 79 sec) [ -23106.9144] [ 4698] [depth= 1] . ( 80 sec) [ -23104.3563] [ 4698] [depth= 4] . ( 80 sec) [ -23101.4684] [ 4696] [depth= 5] . ( 80 sec) [ -23101.2333] [ 4695] [depth= 1] . ( 81 sec) [ -23098.9468] [ 4691] [depth= 1] . ( 81 sec) [ -23076.2103] [ 4690] [depth= 1] . ( 81 sec) [ -23005.0460] [ 4706] [depth= 1] . ( 82 sec) [ -22789.2158] [ 4634] [depth= 5] . ( 83 sec) [ -22757.8464] [ 4626] [depth= 7] . ( 83 sec) [ -22756.9032] [ 4627] [depth= 2] . ( 83 sec) [ -22720.4898] [ 4623] [depth= 10] . ( 83 sec) [ -22710.9983] [ 4622] [depth= 3] . ( 83 sec) [ -22709.2570] [ 4622] [depth= 1] . ( 84 sec) [ -22703.6851] [ 4619] [depth= 8] . ( 84 sec) [ -22699.2695] [ 4618] [depth= 1] . ( 84 sec) [ -22698.5591] [ 4618] [depth= 1] . ( 84 sec) [ -22696.9899] [ 4618] [depth= 1] . ( 85 sec) [ -22696.7893] [ 4618] [depth= 1] . ( 85 sec) [ -22606.0620] [ 4588] [depth= 1] . ( 85 sec) [ -22604.1222] [ 4596] [depth= 1] . ( 86 sec) [ -22603.8901] [ 4597] [depth= 3] . ( 86 sec) [ -22601.5976] [ 4594] [depth= 7] . ( 86 sec) [ -22592.3481] [ 4600] [depth= 1] . ( 86 sec) [ -22592.1538] [ 4601] [depth= 1] . ( 87 sec) [ -22589.8863] [ 4600] [depth= 1] . ( 87 sec) [ -22565.1163] [ 4594] [depth= 1] . ( 87 sec) [ -22519.4511] [ 4571] [depth= 9] . ( 88 sec) [ -22497.9729] [ 4566] [depth= 7] . ( 88 sec) [ -22493.3924] [ 4565] [depth= 1] . ( 89 sec) [ -22459.4273] [ 4557] [depth= 8] . ( 89 sec) [ -22454.9192] [ 4557] [depth= 1] . ( 89 sec) [ -22444.6934] [ 4554] [depth= 2] . ( 90 sec) [ -22444.2863] [ 4556] [depth= 1] . ( 90 sec) [ -22435.7274] [ 4555] [depth= 1] . ( 90 sec) [ -22433.7815] [ 4554] [depth= 1] . ( 91 sec) [ -22400.1906] [ 4554] [depth= 1] . ( 91 sec) [ -22388.7265] [ 4555] [depth= 1] . ( 91 sec) [ -22387.2868] [ 4553] [depth= 2] . ( 92 sec) [ -22382.1733] [ 4549] [depth= 11] . ( 92 sec) [ -22378.4428] [ 4553] [depth= 12] . ( 92 sec) [ -22376.1745] [ 4550] [depth= 1] . ( 93 sec) [ -22370.5868] [ 4548] [depth= 7] . ( 93 sec) [ -22360.2505] [ 4543] [depth= 7] . ( 93 sec) [ -22350.2254] [ 4541] [depth= 2] . ( 94 sec) [ -22343.1480] [ 4540] [depth= 1] . ( 94 sec) [ -22341.7452] [ 4540] [depth= 1] . ( 94 sec) [ -22336.8270] [ 4541] [depth= 1] . ( 94 sec) [ -22316.5027] [ 4533] [depth= 6] . ( 95 sec) [ -22311.3050] [ 4535] [depth= 4] . ( 95 sec) [ -22250.2839] [ 4538] [depth= 11] . ( 95 sec) [ -22210.5454] [ 4532] [depth= 5] . ( 96 sec) [ -22209.5319] [ 4532] [depth= 1] . ( 98 sec) [ -21731.4567] [Branch lengths ] . ( 105 sec) [ -21711.1455] [GTR parameters ] . ( 106 sec) [ -21643.7359] [Alpha ][ 0.332772] . ( 106 sec) [ -21643.7124] [ 4532] [depth= 1] . ( 107 sec) [ -21636.4191] [ 4531] [depth= 1] . ( 108 sec) [ -21636.4087] [ 4532] [depth= 1] . ( 108 sec) [ -21629.7929] [ 4530] [depth= 1] . ( 110 sec) [ -21629.2121] [ 4529] [depth= 1] . ( 111 sec) [ -21613.9055] [ 4521] [depth= 1] . ( 112 sec) [ -21611.0586] [ 4521] [depth= 2] . ( 112 sec) [ -21602.3366] [ 4519] [depth= 4] . ( 112 sec) [ -21596.5254] [ 4515] [depth= 1] . ( 113 sec) [ -21564.5438] [ 4491] [depth= 17] . ( 113 sec) [ -21564.3509] [ 4492] [depth= 10] . ( 114 sec) [ -21538.0648] [ 4482] [depth= 1] . ( 114 sec) [ -21522.0103] [ 4464] [depth= 9] . ( 115 sec) [ -21507.9110] [ 4462] [depth= 3] . ( 116 sec) [ -21476.4450] [ 4449] [depth= 11] . ( 116 sec) [ -21465.3054] [ 4450] [depth= 1] . ( 116 sec) [ -21460.0186] [ 4446] [depth= 1] . ( 117 sec) [ -21319.3048] [ 4371] [depth= 1] . ( 117 sec) [ -21243.2991] [ 4331] [depth= 9] . ( 119 sec) [ -21235.2803] [ 4316] [depth= 1] . ( 122 sec) [ -21226.0619] [ 4314] [depth= 1] . ( 122 sec) [ -21225.9128] [ 4315] [depth= 1] . ( 123 sec) [ -21225.2758] [ 4314] [depth= 1] . ( 123 sec) [ -21224.8239] [ 4314] [depth= 1] . ( 123 sec) [ -21224.1602] [ 4313] [depth= 2] . ( 124 sec) [ -21223.0890] [ 4314] [depth= 1] . ( 125 sec) [ -21220.8126] [ 4310] [depth= 4] . ( 127 sec) [ -21217.1960] [ 4309] [depth= 1] . ( 128 sec) [ -21216.1362] [ 4290] [depth= 1] . ( 129 sec) [ -21214.6206] [ 4288] [depth= 3] . ( 133 sec) [ -21211.0360] [ 4286] [depth= 1] . ( 134 sec) [ -21210.1194] [ 4284] [depth= 8] . ( 134 sec) [ -21207.2303] [ 4285] [depth= 1] . ( 134 sec) [ -21196.1299] [ 4283] [depth= 1] . ( 135 sec) [ -21193.7699] [ 4274] [depth= 5] . ( 135 sec) [ -21081.8640] [ 4241] [depth= 1] . ( 135 sec) [ -21080.7839] [ 4241] [depth= 1] . ( 136 sec) [ -21071.6075] [ 4238] [depth= 3] . ( 136 sec) [ -21071.4684] [ 4238] [depth= 1] . ( 136 sec) [ -21068.2579] [ 4239] [depth= 1] . ( 136 sec) [ -21067.5132] [ 4239] [depth= 1] . ( 137 sec) [ -21060.5637] [ 4236] [depth= 3] . ( 137 sec) [ -21032.1838] [ 4229] [depth= 6] . ( 138 sec) [ -21016.5061] [ 4221] [depth= 6] . ( 139 sec) [ -21016.0801] [ 4221] [depth= 1] . ( 139 sec) [ -21009.0232] [ 4220] [depth= 2] . ( 140 sec) [ -21006.9440] [ 4219] [depth= 3] . ( 140 sec) [ -21001.4391] [ 4218] [depth= 2] . ( 141 sec) [ -20974.1504] [ 4202] [depth= 5] . ( 142 sec) [ -20965.2360] [ 4201] [depth= 2] . ( 142 sec) [ -20924.5898] [ 4186] [depth= 5] . ( 143 sec) [ -20923.7101] [ 4185] [depth= 1] . ( 143 sec) [ -20915.9568] [ 4183] [depth= 3] . ( 145 sec) [ -20911.8705] [ 4180] [depth= 8] . ( 145 sec) [ -20906.1576] [ 4175] [depth= 2] . ( 146 sec) [ -20898.3153] [ 4176] [depth= 1] . ( 146 sec) [ -20898.0255] [ 4176] [depth= 1] . ( 147 sec) [ -20887.8778] [ 4171] [depth= 1] . ( 147 sec) [ -20887.5802] [ 4172] [depth= 1] . ( 149 sec) [ -20887.4377] [ 4172] [depth= 1] . ( 150 sec) [ -20887.0514] [ 4172] [depth= 1] . ( 151 sec) [ -20860.5893] [ 4158] [depth= 2] . ( 153 sec) [ -20832.6591] [ 4148] [depth= 1] . ( 153 sec) [ -20823.9336] [ 4148] [depth= 4] . ( 154 sec) [ -20822.9968] [ 4149] [depth= 1] . ( 154 sec) [ -20821.7838] [ 4148] [depth= 1] . ( 154 sec) [ -20821.1215] [ 4149] [depth= 1] . ( 154 sec) [ -20820.9443] [ 4149] [depth= 1] . ( 155 sec) [ -20819.7309] [ 4147] [depth= 2] . ( 155 sec) [ -20809.4548] [ 4142] [depth= 1] . ( 157 sec) [ -20805.2955] [ 4141] [depth= 5] . ( 157 sec) [ -20804.8621] [ 4141] [depth= 1] . ( 158 sec) [ -20804.2959] [ 4141] [depth= 1] . ( 158 sec) [ -20804.1256] [ 4141] [depth= 1] . ( 158 sec) [ -20802.2534] [ 4137] [depth= 1] . ( 159 sec) [ -20798.6653] [ 4136] [depth= 1] . ( 160 sec) [ -20796.8940] [ 4135] [depth= 4] . ( 160 sec) [ -20783.1803] [ 4132] [depth= 2] . ( 161 sec) [ -20781.9473] [ 4132] [depth= 1] . ( 161 sec) [ -20779.0066] [ 4130] [depth= 8] . ( 162 sec) [ -20774.2814] [ 4131] [depth= 1] . ( 162 sec) [ -20770.3105] [ 4128] [depth= 1] . ( 163 sec) [ -20747.1143] [ 4118] [depth= 9] . ( 163 sec) [ -20735.6329] [ 4112] [depth= 8] . ( 164 sec) [ -20732.1113] [ 4112] [depth= 2] . ( 165 sec) [ -20731.3721] [ 4113] [depth= 1] . ( 166 sec) [ -20731.1960] [ 4115] [depth= 7] . ( 166 sec) [ -20728.1733] [ 4114] [depth= 1] . ( 169 sec) [ -20718.3642] [ 4109] [depth= 1] . ( 169 sec) [ -20717.5830] [ 4108] [depth= 1] . ( 170 sec) [ -20712.2859] [ 4097] [depth= 8] . ( 171 sec) [ -20707.0875] [ 4095] [depth= 1] . ( 172 sec) [ -20702.3370] [ 4090] [depth= 7] . ( 172 sec) [ -20701.1439] [ 4091] [depth= 1] . ( 173 sec) [ -20698.0895] [ 4089] [depth= 1] . ( 173 sec) [ -20681.3077] [ 4082] [depth= 7] . ( 173 sec) [ -20664.1316] [ 4071] [depth= 1] . ( 173 sec) [ -20662.0470] [ 4071] [depth= 1] . ( 174 sec) [ -20660.3085] [ 4075] [depth= 1] . ( 174 sec) [ -20657.0602] [ 4071] [depth= 6] . ( 175 sec) [ -20656.2340] [ 4070] [depth= 2] . ( 176 sec) [ -20654.4286] [ 4065] [depth= 1] . ( 178 sec) [ -20653.9561] [ 4067] [depth= 1] . ( 179 sec) [ -20653.8124] [ 4067] [depth= 1] . ( 180 sec) [ -20652.6366] [ 4065] [depth= 1] . ( 180 sec) [ -20649.8192] [ 4064] [depth= 1] . ( 184 sec) [ -20639.4950] [ 4060] [depth= 1] . ( 184 sec) [ -20636.5514] [ 4051] [depth= 3] . ( 185 sec) [ -20631.5808] [ 4050] [depth= 1] . ( 186 sec) [ -20627.3695] [ 4051] [depth= 1] . ( 188 sec) [ -20626.8850] [ 4051] [depth= 1] . ( 188 sec) [ -20626.4326] [ 4052] [depth= 1] . ( 190 sec) [ -20597.9252] [ 4051] [depth= 4] . ( 191 sec) [ -20597.5317] [ 4053] [depth= 1] . ( 193 sec) [ -20597.4210] [ 4052] [depth= 1] . ( 195 sec) [ -20594.4783] [ 4054] [depth= 1] . ( 195 sec) [ -20593.0610] [ 4053] [depth= 1] . ( 196 sec) [ -20586.5441] [ 4051] [depth= 2] . ( 197 sec) [ -20586.5078] [ 4051] [depth= 1] . ( 197 sec) [ -20570.5501] [ 4048] [depth= 1] . ( 198 sec) [ -20569.9007] [ 4047] [depth= 1] . ( 199 sec) [ -20567.7177] [ 4047] [depth= 2] . ( 199 sec) [ -20565.7256] [ 4047] [depth= 1] . ( 199 sec) [ -20564.1872] [ 4049] [depth= 1] . ( 202 sec) [ -20563.9713] [ 4049] [depth= 1] . ( 203 sec) [ -20563.5222] [ 4049] [depth= 1] . ( 204 sec) [ -20561.9169] [ 4048] [depth= 1] . ( 204 sec) [ -20561.1582] [ 4049] [depth= 2] . ( 205 sec) [ -20560.9977] [ 4049] [depth= 1] . ( 206 sec) [ -20560.1736] [ 4051] [depth= 2] . ( 206 sec) [ -20549.3623] [ 4045] [depth= 4] . ( 207 sec) [ -20548.9907] [ 4046] [depth= 1] . ( 210 sec) [ -20465.3868] [Branch lengths ] . ( 214 sec) [ -20463.3184] [GTR parameters ] . ( 214 sec) [ -20463.2175] [Alpha ][ 0.337974] . ( 215 sec) [ -20463.2049] [ 4046] [depth= 1] . ( 216 sec) [ -20461.7592] [ 4044] [depth= 2] . ( 228 sec) [ -20460.0091] [ 4041] [depth= 3] . ( 231 sec) [ -20459.1167] [ 4042] [depth= 1] . ( 234 sec) [ -20458.9711] [ 4042] [depth= 1] . ( 238 sec) [ -20458.4959] [ 4045] [depth= 4] . ( 245 sec) [ -20457.7632] [ 4046] [depth= 2] . ( 247 sec) [ -20452.6304] [ 4043] [depth= 7] . ( 247 sec) [ -20452.5456] [ 4042] [depth= 1] . ( 252 sec) [ -20452.2432] [ 4042] [depth= 1] . ( 254 sec) [ -20451.4908] [ 4043] [depth= 1] . ( 260 sec) [ -20443.6069] [ 4040] [depth= 2] . ( 265 sec) [ -20442.7083] [ 4038] [depth= 1] . ( 268 sec) [ -20442.4213] [ 4040] [depth= 1] . ( 271 sec) [ -20434.1190] [ 4037] [depth= 1] . ( 272 sec) [ -20433.0370] [ 4036] [depth= 1] . ( 272 sec) [ -20431.2391] [ 4031] [depth= 1] . ( 275 sec) [ -20430.8739] [ 4031] [depth= 1] . ( 287 sec) [ -20430.8519] [ 4035] [depth= 2] . ( 288 sec) [ -20428.6560] [ 4033] [depth= 2] . ( 291 sec) [ -20427.2419] [ 4036] [depth= 1] . ( 302 sec) [ -20426.9327] [ 4035] [depth= 1] . ( 304 sec) [ -20426.8328] [ 4035] [depth= 1] . ( 305 sec) [ -20426.7461] [ 4036] [depth= 1] . ( 311 sec) [ -20425.3230] [ 4037] [depth= 1] . ( 314 sec) [ -20425.2882] [ 4037] [depth= 1] . ( 331 sec) [ -20425.2168] [ 4039] [depth= 1] . ( 343 sec) [ -20415.0515] [Branch lengths ] . ( 347 sec) [ -20414.9634] [GTR parameters ] . ( 348 sec) [ -20414.9588] [Alpha ][ 0.339038] . ( 353 sec) [ -20412.0749] [ 4036] [depth= 9] . ( 358 sec) [ -20412.0177] [ 4034] [depth= 1] . ( 358 sec) [ -20411.5425] [ 4033] [depth= 1] . ( 361 sec) [ -20410.6185] [ 4037] [depth= 1] . ( 369 sec) [ -20410.2175] [ 4036] [depth= 1] . ( 376 sec) [ -20409.7627] [ 4035] [depth= 1] . ( 390 sec) [ -20409.6602] [ 4035] [depth= 1] . ( 393 sec) [ -20409.0586] [ 4035] [depth= 1] . ( 404 sec) [ -20408.9135] [ 4033] [depth= 1] . ( 415 sec) [ -20408.9018] [ 4032] [depth= 1] . ( 436 sec) [ -20408.7837] [ 4033] [depth= 1] . ( 448 sec) [ -20408.7482] [ 4032] [depth= 1] . ( 471 sec) [ -20403.4755] [Branch lengths ] . ( 475 sec) [ -20403.4049] [GTR parameters ] . ( 475 sec) [ -20403.3979] [Alpha ][ 0.340365] . ( 600 sec) [ -20402.9734] [Branch lengths ] . ( 792 sec) [ -20402.7493] [ 4048] [depth= 1] . ( 851 sec) [ -20402.5821] [Branch lengths ] . ( 854 sec) [ -20402.5753] [GTR parameters ] . ( 855 sec) [ -20402.5727] [Alpha ][ 0.341155] . ( 855 sec) [ -20402.5727] [Topology ] . ( 860 sec) [ -20402.3178] [Topology ] . ( 865 sec) [ -20402.2737] [Topology ] . ( 871 sec) [ -20402.2654] [Topology ] . ( 876 sec) [ -20402.2630] [Topology ] . ( 881 sec) [ -20402.2623] [Topology ] . ( 882 sec) [ -20402.2620] [GTR parameters ] . ( 883 sec) [ -20402.2620] [Alpha ][ 0.341155] . ( 883 sec) [ -20402.2620] [Topology ] . ( 888 sec) [ -20402.2617] [Topology ] . ( 889 sec) [ -20402.2617] [GTR parameters ] . ( 889 sec) [ -20402.2617] [Alpha ][ 0.341155] . Checking for NNIs, optimizing five branches... . ( 927 sec) [ -20401.6359] [Topology ] . Checking for NNIs, optimizing five branches... . ( 964 sec) [ -20401.1928] [Topology ] . Checking for NNIs, optimizing five branches... . ( 1001 sec) [ -20401.1081] [Topology ] . Checking for NNIs, optimizing five branches... . Log likelihood of the current tree: -20401.108102. . Computing pairwise distances... . Building BioNJ tree... . Maximizing likelihood (using NNI moves)... . ( 7 sec) [ -21575.3354] [GTR parameters ] . ( 8 sec) [ -21206.7214] [Alpha ][ 0.500960] . ( 8 sec) [ -21206.7214] [Topology ] . ( 13 sec) [ -20737.3169] [Topology ] . ( 18 sec) [ -20622.9577] [Topology ] . ( 23 sec) [ -20586.6538] [Topology ] . ( 28 sec) [ -20570.9427] [Topology ] . ( 34 sec) [ -20566.3417] [Topology ] . ( 39 sec) [ -20561.9455] [Topology ] . ( 44 sec) [ -20559.4520] [Topology ] . ( 49 sec) [ -20551.4915] [Topology ] . ( 54 sec) [ -20546.5863] [Topology ] . Checking for NNIs, optimizing five branches... . ( 98 sec) [ -20541.1422] [Topology ] . ( 103 sec) [ -20529.9550] [GTR parameters ] . ( 103 sec) [ -20478.5225] [Alpha ][ 0.348656] . ( 103 sec) [ -20478.5225] [Topology ] . ( 109 sec) [ -20471.3446] [Topology ] . ( 114 sec) [ -20470.4270] [Topology ] . ( 119 sec) [ -20469.8076] [Topology ] . ( 124 sec) [ -20469.2380] [Topology ] . ( 129 sec) [ -20469.1369] [Topology ] . ( 135 sec) [ -20469.1145] [Topology ] . ( 140 sec) [ -20469.1090] [Topology ] . ( 145 sec) [ -20469.1075] [Topology ] . ( 150 sec) [ -20469.1071] [Topology ] . Checking for NNIs, optimizing five branches... . ( 187 sec) [ -20468.5217] [Topology ] . ( 190 sec) [ -20468.0511] [GTR parameters ] . ( 191 sec) [ -20467.8627] [Alpha ][ 0.341568] . ( 191 sec) [ -20467.8627] [Topology ] . ( 196 sec) [ -20467.5304] [Topology ] . ( 201 sec) [ -20467.1917] [Topology ] . ( 206 sec) [ -20465.9457] [Topology ] . ( 211 sec) [ -20465.8018] [Topology ] . ( 216 sec) [ -20465.7725] [Topology ] . ( 222 sec) [ -20465.7638] [Topology ] . ( 227 sec) [ -20465.7610] [Topology ] . ( 232 sec) [ -20465.7599] [Topology ] . Checking for NNIs, optimizing five branches... . ( 269 sec) [ -20465.2554] [Topology ] . ( 271 sec) [ -20465.2491] [GTR parameters ] . ( 271 sec) [ -20465.2471] [Alpha ][ 0.340728] . ( 271 sec) [ -20465.2471] [Topology ] . ( 276 sec) [ -20465.2386] [Topology ] . ( 282 sec) [ -20465.2368] [Topology ] . ( 287 sec) [ -20465.2364] [Topology ] . Checking for NNIs, optimizing five branches... . ( 324 sec) [ -20464.7322] [Topology ] . ( 326 sec) [ -20464.7257] [GTR parameters ] . ( 326 sec) [ -20464.7251] [Alpha ][ 0.341069] . ( 326 sec) [ -20464.7251] [Topology ] . ( 331 sec) [ -20464.6624] [Topology ] . ( 336 sec) [ -20464.6484] [Topology ] . ( 341 sec) [ -20464.6430] [Topology ] . ( 347 sec) [ -20464.6405] [Topology ] . ( 352 sec) [ -20464.6392] [Topology ] . ( 357 sec) [ -20464.6386] [Topology ] . Checking for NNIs, optimizing five branches... . ( 394 sec) [ -20464.6349] [Topology ] . ( 395 sec) [ -20464.6341] [GTR parameters ] . ( 395 sec) [ -20464.6335] [Alpha ][ 0.341410] . ( 395 sec) [ -20464.6335] [Topology ] . ( 400 sec) [ -20464.6329] [Topology ] . Checking for NNIs, optimizing five branches... . ( 438 sec) [ -20464.6329] [GTR parameters ] . ( 438 sec) [ -20464.6329] [Alpha ][ 0.341410] . ( 439 sec) [ -20464.6329] [Topology ] . ( 444 sec) [ -20464.6326] [Topology ] . Checking for NNIs, optimizing five branches... . Checking for NNIs, optimizing five branches... . Maximizing likelihood (using SPR moves)... . ( 524 sec) [ -21575.3354] [GTR parameters ] . ( 524 sec) [ -21206.7214] [Alpha ][ 0.500960] . ( 525 sec) [ -21206.1411] [ 4073] [depth= 1] . ( 526 sec) [ -21199.5150] [ 4077] [depth= 1] . ( 526 sec) [ -21185.9218] [ 4073] [depth= 1] . ( 526 sec) [ -21171.3223] [ 4059] [depth= 1] . ( 527 sec) [ -21169.7149] [ 4065] [depth= 1] . ( 528 sec) [ -21169.1018] [ 4076] [depth= 1] . ( 528 sec) [ -21148.9782] [ 4065] [depth= 1] . ( 529 sec) [ -21148.1301] [ 4070] [depth= 1] . ( 529 sec) [ -21138.2511] [ 4065] [depth= 1] . ( 530 sec) [ -21132.9040] [ 4069] [depth= 1] . ( 530 sec) [ -21118.3850] [ 4065] [depth= 1] . ( 531 sec) [ -21116.7828] [ 4067] [depth= 1] . ( 531 sec) [ -21107.9046] [ 4065] [depth= 1] . ( 532 sec) [ -21107.6878] [ 4069] [depth= 1] . ( 532 sec) [ -21102.5459] [ 4081] [depth= 1] . ( 532 sec) [ -21073.5167] [ 4069] [depth= 1] . ( 533 sec) [ -21067.2112] [ 4065] [depth= 1] . ( 534 sec) [ -21042.3328] [ 4074] [depth= 4] . ( 534 sec) [ -21031.7104] [ 4078] [depth= 1] . ( 534 sec) [ -21031.5153] [ 4074] [depth= 1] . ( 535 sec) [ -21031.0870] [ 4070] [depth= 1] . ( 535 sec) [ -21030.5398] [ 4070] [depth= 2] . ( 536 sec) [ -21030.4456] [ 4074] [depth= 1] . ( 536 sec) [ -21029.3591] [ 4077] [depth= 1] . ( 536 sec) [ -21028.4497] [ 4078] [depth= 1] . ( 537 sec) [ -21027.1816] [ 4074] [depth= 1] . ( 537 sec) [ -21026.3218] [ 4077] [depth= 1] . ( 537 sec) [ -21024.7836] [ 4074] [depth= 1] . ( 538 sec) [ -21024.5493] [ 4078] [depth= 1] . ( 538 sec) [ -21016.1002] [ 4080] [depth= 1] . ( 539 sec) [ -21012.7409] [ 4079] [depth= 1] . ( 539 sec) [ -21008.0453] [ 4080] [depth= 1] . ( 540 sec) [ -20998.6533] [ 4077] [depth= 1] . ( 540 sec) [ -20997.7683] [ 4078] [depth= 1] . ( 541 sec) [ -20996.9510] [ 4077] [depth= 1] . ( 541 sec) [ -20991.8876] [ 4076] [depth= 1] . ( 541 sec) [ -20986.3212] [ 4077] [depth= 1] . ( 542 sec) [ -20982.1182] [ 4078] [depth= 1] . ( 544 sec) [ -20981.9009] [ 4079] [depth= 1] . ( 544 sec) [ -20981.6467] [ 4078] [depth= 3] . ( 544 sec) [ -20981.1407] [ 4079] [depth= 1] . ( 545 sec) [ -20953.4563] [ 4073] [depth= 4] . ( 545 sec) [ -20947.8959] [ 4071] [depth= 2] . ( 547 sec) [ -20946.9900] [ 4073] [depth= 1] . ( 547 sec) [ -20944.4483] [ 4071] [depth= 1] . ( 547 sec) [ -20943.7214] [ 4071] [depth= 1] . ( 548 sec) [ -20943.4590] [ 4072] [depth= 1] . ( 548 sec) [ -20941.0003] [ 4068] [depth= 12] . ( 549 sec) [ -20932.6285] [ 4067] [depth= 6] . ( 550 sec) [ -20924.4043] [ 4066] [depth= 2] . ( 550 sec) [ -20920.8819] [ 4064] [depth= 7] . ( 550 sec) [ -20919.2612] [ 4064] [depth= 2] . ( 551 sec) [ -20918.2796] [ 4064] [depth= 1] . ( 551 sec) [ -20897.8781] [ 4063] [depth= 2] . ( 551 sec) [ -20892.2793] [ 4056] [depth= 5] . ( 551 sec) [ -20890.2083] [ 4056] [depth= 2] . ( 552 sec) [ -20889.8978] [ 4056] [depth= 1] . ( 552 sec) [ -20889.7377] [ 4056] [depth= 1] . ( 553 sec) [ -20885.1230] [ 4057] [depth= 2] . ( 553 sec) [ -20883.7516] [ 4057] [depth= 1] . ( 554 sec) [ -20883.6787] [ 4062] [depth= 1] . ( 555 sec) [ -20877.6561] [ 4063] [depth= 1] . ( 555 sec) [ -20877.1934] [ 4062] [depth= 1] . ( 555 sec) [ -20876.0042] [ 4062] [depth= 1] . ( 556 sec) [ -20875.3066] [ 4062] [depth= 1] . ( 556 sec) [ -20874.4625] [ 4062] [depth= 1] . ( 556 sec) [ -20867.5433] [ 4061] [depth= 6] . ( 557 sec) [ -20867.5068] [ 4061] [depth= 1] . ( 557 sec) [ -20866.3044] [ 4060] [depth= 1] . ( 559 sec) [ -20864.0107] [ 4059] [depth= 2] . ( 559 sec) [ -20863.9519] [ 4063] [depth= 1] . ( 559 sec) [ -20863.1543] [ 4059] [depth= 1] . ( 560 sec) [ -20861.3639] [ 4060] [depth= 1] . ( 562 sec) [ -20861.1209] [ 4061] [depth= 2] . ( 564 sec) [ -20859.3721] [ 4061] [depth= 2] . ( 565 sec) [ -20858.6604] [ 4061] [depth= 1] . ( 566 sec) [ -20857.7099] [ 4061] [depth= 1] . ( 566 sec) [ -20857.4707] [ 4061] [depth= 1] . ( 568 sec) [ -20857.4052] [ 4061] [depth= 1] . ( 568 sec) [ -20856.7512] [ 4062] [depth= 1] . ( 568 sec) [ -20856.3820] [ 4062] [depth= 1] . ( 569 sec) [ -20855.4101] [ 4062] [depth= 1] . ( 569 sec) [ -20853.0166] [ 4062] [depth= 1] . ( 571 sec) [ -20851.1565] [ 4064] [depth= 1] . ( 571 sec) [ -20844.2644] [ 4062] [depth= 1] . ( 572 sec) [ -20844.2112] [ 4062] [depth= 1] . ( 572 sec) [ -20844.0141] [ 4062] [depth= 1] . ( 572 sec) [ -20843.7250] [ 4062] [depth= 1] . ( 572 sec) [ -20843.2067] [ 4062] [depth= 2] . ( 573 sec) [ -20837.7391] [ 4061] [depth= 1] . ( 573 sec) [ -20832.8016] [ 4060] [depth= 1] . ( 573 sec) [ -20832.0503] [ 4059] [depth= 1] . ( 574 sec) [ -20828.0222] [ 4059] [depth= 1] . ( 575 sec) [ -20827.5277] [ 4060] [depth= 1] . ( 577 sec) [ -20815.1555] [ 4055] [depth= 7] . ( 578 sec) [ -20809.7885] [ 4055] [depth= 1] . ( 578 sec) [ -20807.5134] [ 4055] [depth= 1] . ( 579 sec) [ -20804.0305] [ 4054] [depth= 1] . ( 580 sec) [ -20800.0927] [ 4054] [depth= 1] . ( 581 sec) [ -20799.8763] [ 4057] [depth= 1] . ( 583 sec) [ -20796.6724] [ 4057] [depth= 1] . ( 584 sec) [ -20793.7520] [ 4056] [depth= 1] . ( 587 sec) [ -20789.9325] [ 4060] [depth= 4] . ( 587 sec) [ -20787.1378] [ 4056] [depth= 4] . ( 587 sec) [ -20775.3477] [ 4053] [depth= 7] . ( 588 sec) [ -20773.9425] [ 4053] [depth= 1] . ( 588 sec) [ -20772.5686] [ 4054] [depth= 2] . ( 589 sec) [ -20770.5859] [ 4053] [depth= 1] . ( 590 sec) [ -20770.2396] [ 4055] [depth= 8] . ( 590 sec) [ -20769.4393] [ 4053] [depth= 6] . ( 595 sec) [ -20766.0015] [ 4052] [depth= 2] . ( 595 sec) [ -20765.5721] [ 4051] [depth= 1] . ( 596 sec) [ -20760.5993] [ 4048] [depth= 1] . ( 600 sec) [ -20760.3302] [ 4051] [depth= 2] . ( 601 sec) [ -20758.9319] [ 4050] [depth= 3] . ( 601 sec) [ -20757.6391] [ 4050] [depth= 2] . ( 602 sec) [ -20745.2456] [ 4045] [depth= 3] . ( 602 sec) [ -20745.0911] [ 4047] [depth= 2] . ( 602 sec) [ -20743.1663] [ 4045] [depth= 2] . ( 603 sec) [ -20742.9972] [ 4046] [depth= 1] . ( 603 sec) [ -20742.6183] [ 4045] [depth= 1] . ( 603 sec) [ -20742.5467] [ 4045] [depth= 1] . ( 605 sec) [ -20741.4109] [ 4045] [depth= 1] . ( 606 sec) [ -20741.2551] [ 4045] [depth= 1] . ( 606 sec) [ -20739.1957] [ 4045] [depth= 1] . ( 606 sec) [ -20738.7493] [ 4045] [depth= 1] . ( 608 sec) [ -20738.3673] [ 4045] [depth= 1] . ( 609 sec) [ -20737.1109] [ 4044] [depth= 4] . ( 609 sec) [ -20737.0004] [ 4044] [depth= 2] . ( 609 sec) [ -20735.0054] [ 4044] [depth= 1] . ( 615 sec) [ -20734.8551] [ 4045] [depth= 1] . ( 616 sec) [ -20732.5964] [ 4045] [depth= 1] . ( 617 sec) [ -20730.3801] [ 4045] [depth= 2] . ( 618 sec) [ -20730.1478] [ 4045] [depth= 1] . ( 619 sec) [ -20729.3912] [ 4045] [depth= 1] . ( 620 sec) [ -20728.9981] [ 4046] [depth= 1] . ( 623 sec) [ -20727.1255] [ 4044] [depth= 1] . ( 625 sec) [ -20553.4281] [Branch lengths ] . ( 630 sec) [ -20545.2582] [GTR parameters ] . ( 631 sec) [ -20488.8294] [Alpha ][ 0.339831] . ( 631 sec) [ -20486.0947] [ 4044] [depth= 1] . ( 632 sec) [ -20484.0219] [ 4050] [depth= 1] . ( 639 sec) [ -20480.7496] [ 4052] [depth= 5] . ( 639 sec) [ -20480.3805] [ 4052] [depth= 1] . ( 641 sec) [ -20480.0668] [ 4049] [depth= 1] . ( 642 sec) [ -20473.9225] [ 4043] [depth= 1] . ( 643 sec) [ -20473.0011] [ 4047] [depth= 1] . ( 651 sec) [ -20472.8530] [ 4047] [depth= 1] . ( 660 sec) [ -20472.6913] [ 4048] [depth= 1] . ( 662 sec) [ -20472.6650] [ 4048] [depth= 1] . ( 665 sec) [ -20472.6147] [ 4049] [depth= 1] . ( 666 sec) [ -20472.5287] [ 4048] [depth= 1] . ( 673 sec) [ -20472.4525] [ 4048] [depth= 1] . ( 675 sec) [ -20472.4280] [ 4048] [depth= 1] . ( 679 sec) [ -20470.0690] [ 4047] [depth= 2] . ( 681 sec) [ -20469.0941] [ 4047] [depth= 1] . ( 683 sec) [ -20468.9624] [ 4047] [depth= 1] . ( 684 sec) [ -20462.6495] [ 4044] [depth= 1] . ( 684 sec) [ -20461.8610] [ 4044] [depth= 1] . ( 695 sec) [ -20460.6745] [ 4045] [depth= 3] . ( 700 sec) [ -20459.6244] [ 4046] [depth= 7] . ( 704 sec) [ -20457.3936] [ 4046] [depth= 1] . ( 705 sec) [ -20457.0632] [ 4046] [depth= 1] . ( 705 sec) [ -20455.9182] [ 4046] [depth= 1] . ( 705 sec) [ -20455.6620] [ 4046] [depth= 1] . ( 715 sec) [ -20455.4574] [ 4047] [depth= 2] . ( 720 sec) [ -20454.6221] [ 4049] [depth= 7] . ( 721 sec) [ -20454.2434] [ 4048] [depth= 1] . ( 723 sec) [ -20454.2287] [ 4048] [depth= 1] . ( 725 sec) [ -20454.1110] [ 4048] [depth= 1] . ( 726 sec) [ -20453.6661] [ 4049] [depth= 1] . ( 726 sec) [ -20453.0362] [ 4047] [depth= 1] . ( 737 sec) [ -20452.7598] [ 4051] [depth= 7] . ( 738 sec) [ -20451.9105] [ 4050] [depth= 1] . ( 738 sec) [ -20451.6577] [ 4052] [depth= 1] . ( 738 sec) [ -20450.5604] [ 4049] [depth= 2] . ( 742 sec) [ -20422.2990] [Branch lengths ] . ( 746 sec) [ -20421.1370] [GTR parameters ] . ( 747 sec) [ -20421.1312] [Alpha ][ 0.338476] . ( 753 sec) [ -20420.7962] [ 4053] [depth= 1] . ( 756 sec) [ -20419.2140] [ 4053] [depth= 1] . ( 770 sec) [ -20418.8796] [ 4057] [depth= 15] . ( 773 sec) [ -20418.8638] [ 4057] [depth= 1] . ( 774 sec) [ -20418.8467] [ 4057] [depth= 1] . ( 784 sec) [ -20418.7989] [ 4058] [depth= 1] . ( 814 sec) [ -20417.8336] [ 4057] [depth= 1] . ( 855 sec) [ -20417.6036] [ 4057] [depth= 1] . ( 889 sec) [ -20413.7598] [Branch lengths ] . ( 892 sec) [ -20413.6812] [GTR parameters ] . ( 893 sec) [ -20413.6753] [Alpha ][ 0.339677] . ( 902 sec) [ -20413.0862] [ 4059] [depth= 3] . ( 918 sec) [ -20413.0575] [ 4061] [depth= 2] . ( 923 sec) [ -20411.9624] [ 4059] [depth= 3] . ( 926 sec) [ -20410.8771] [ 4061] [depth= 1] . ( 926 sec) [ -20410.8539] [ 4062] [depth= 1] . ( 926 sec) [ -20410.3091] [ 4062] [depth= 1] . ( 932 sec) [ -20410.2517] [ 4061] [depth= 1] . ( 1039 sec) [ -20409.0614] [Branch lengths ] . ( 1042 sec) [ -20409.0349] [GTR parameters ] . ( 1042 sec) [ -20409.0349] [Alpha ][ 0.339677] . ( 1063 sec) [ -20408.8730] [ 4062] [depth= 1] . ( 1190 sec) [ -20408.0815] [Branch lengths ] . ( 1192 sec) [ -20408.0784] [GTR parameters ] . ( 1192 sec) [ -20408.0780] [Alpha ][ 0.340017] . ( 1397 sec) [ -20408.0531] [ 4061] [depth= 1] . ( 1441 sec) [ -20406.7347] [Branch lengths ] . ( 1442 sec) [ -20406.7334] [GTR parameters ] . ( 1443 sec) [ -20406.7334] [Alpha ][ 0.340017] . ( 1694 sec) [ -20404.3797] [Branch lengths ] . ( 1694 sec) [ -20404.3797] [Topology ] . ( 1699 sec) [ -20401.7950] [Topology ] . ( 1704 sec) [ -20398.3187] [Topology ] . ( 1709 sec) [ -20397.5064] [Topology ] . ( 1714 sec) [ -20397.3467] [Topology ] . ( 1719 sec) [ -20397.3155] [Topology ] . ( 1725 sec) [ -20397.3090] [Topology ] . ( 1726 sec) [ -20397.3079] [GTR parameters ] . ( 1726 sec) [ -20397.3077] [Alpha ][ 0.340357] . ( 1727 sec) [ -20397.3077] [Topology ] . ( 1732 sec) [ -20397.3062] [Topology ] . ( 1737 sec) [ -20397.3057] [Topology ] . ( 1738 sec) [ -20397.3057] [GTR parameters ] . ( 1738 sec) [ -20397.3057] [Alpha ][ 0.340357] . ( 1738 sec) [ -20397.3057] [Topology ] . ( 1743 sec) [ -20397.3056] [Topology ] . ( 1744 sec) [ -20397.3056] [GTR parameters ] . ( 1745 sec) [ -20397.3056] [Alpha ][ 0.340357] . Checking for NNIs, optimizing five branches... . ( 1781 sec) [ -20397.2961] [Topology ] . Checking for NNIs, optimizing five branches... . ( 1817 sec) [ -20397.2949] [Topology ] . Checking for NNIs, optimizing five branches... . Log likelihood obtained after NNI moves : -20464.632592 . Log likelihood obtained after SPR moves : -20397.294920 . Log likelihood of the current tree: -20397.294920. . Printing the most likely tree in file 'nucleic_M2564_142x1130_2006.phy_phyml_tree.txt'... . Best log likelihood : -20395.733581 . Time used 0h30m54s oooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooooo 7857.08user 0.00system 2:11:29elapsed 99%CPU (0avgtext+0avgdata 0maxresident)k 0inputs+0outputs (0major+0minor)pagefaults 0swaps